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Prof. Dr. Jost Borcherding

Curriculum vitae


Study of biology at the University of Bochum, Germany


Study of biology at the University of Cologne, Germany


Diplom in biology from the University of Cologne


Ph. D. in biology from the University of Cologne both degrees on "The gonad development and reproduction of Dreissena polymorpha"


Research scientist at the University of Cologne granted by the Federal Environmental Agency (Germany) with the subject:"Macrozoobenthos species as bioindicators in the River Rhine; mussel Dreissena as ecological indicator and test organism"


Part A: The Dreissena-Monitor - a biological early warning system for continuous monitoring of water quality


Part B: Population dynamic, growth and larval development of the zebra mussel in a big stream"


with respect to "Part A", member of the "Federal Government/ Federal States ProjectGroup [Wirkungstests Rhein]" that focussed on "Development, Testing and Installation of Biotests for Water Monitoring of the River Rhine" and gave recommendations for the German Commission for the Protection of the Rhine against Pollution".

Since 1993

Management of the ecological research station of the Zoological Institute at the University of Cologne in Rees-Grietherbusch (as research scientist), which is located in the floodplain of the Lower Rhine near the border to the Netherlands


Habilitation at the University of Cologne on the ecological adaptations of fishes in the face of a changing environment

Since 2017

Adjunct Professor for Animal Ecology at the University of Cologne, Institute for Zoology


Research interests

My research focuses on perch and invasive Gobiids as model organisms, analysing population dynamics, growth, predator-prey relationships, feeding, behaviour, morphometrics and other ecological characteristics in order to understand their impact on fish communities in the River Rhine and surrounding waters. In addition, we are engaged in management and research on migratory fishes in North-Rhine-Westphalia and actually monitor the impact of technically improved hydroelectric plants on the downstream migration of salmon smolts and silver eel.

Recent publications

Pdf-Download here

Articles in peer reviewed journals


Ingendahl, D., Borcherding, J. ( 2022): Zur Bedeutung der Wasserkraft und ihren Auswirkungen auf die Gewässerökologie am Beispiel von Nordrhein-Westfalen. Wasser und Abfall 01-02, 14-19

Heermann L., Teichert, M.A.K., Halang, J., Borcherding, J. (2022): . Does behaviour contribute to the previously documented invasive success of topmouth gudgeon? J.Appl.Ichthyol. n/a (n/a), 2022

Nogueira Tavares, C., Weitere, M, Borcherding, J., Gerngroß, P., Krenek, S., Worischka, S. and Brauns, M. (2022):  Diet composition and trophic niche differentiation of Neogobius melanostomus along an invasion gradient in a large lowland river. Limnologica 95:125996


Borcherding, J., Aschemeier, D., Bruhy, J., Heermann, L., Lindner, J., Schröder, S. L., Wagner, K., Staas, S. (2021): The Caucasian dwarf goby, a new alien Gobiidae spreading at the Lower Rhine, Germany. J. Appl. Ichthyol. doi/10.1111/jai.14196

Ingendahl, D., Heermann, L., Emde, S., Staas, S., Teichert, M., Borcherding, J. (2021): Aalabwanderung an der Wasserkraftanlage Unkelmühle - zwei Methoden, ein Ergebnis? Wasserwirtschaft 9-10/2021, 62-67

Ondrackova, M. Janác, Borcherding, J., Grabowska, J., Bartakova, V. and Jurajda, P. (2021):  Non-native gobies share predominantly immature parasites with local fish hosts. Journal of Vertebrate Biology 70 (4):21050

Tietler, F., Breiden, S., Didenko, A., Storm, S., Borcherding, J. (2021): Identification key for the early juvenile stages of the most important invasive gobies in European freshwaters –A tool for daily practice. J Appl Ichthyol.37:963–967. DOI: 10.1111/jai.14232

Wolter, C., Borcherding, J., Ferreira, T., Freyhof, J., Gessner, J., Górski, j., Nastase, A., Schomaker, C., Erös, T.. (2021): Characterization of European lampreys and fishes by their longitudinal and lateral distribution traits. Ecological Indicators 123


Borcherding, J., Teichert, M., Heermann, L. (2020): Empfehlungen zum Schutz abwandernder Fische an Wasserkraftanlagen. Wasser u Abfall 09: 36-43

Borcherding, J., Webster, M.M., Heubel, K. (2020): Teaching laboratory for large cohorts of undergraduates: Private and social information in fish. Ecology and Evolution 10 (1): 4-10

Havn, T.B., Thorstad, E.B., Borcherding, J., Heermann, L., Teichert, M.A.K., Ingendahl, D., Tambets, M., Sæther, S. A., Økland, F. (2020): Impacts of a weir and power station on downstream migrating Atlantic salmon smolts in a German river. River Research and Applications 36: 784-796. doi: 10.1002/rra.3590

Hempel, C. A., Jacobs, G., Beermann, A. J., Elbrecht, V., Macher, J.-N., Macher, T.-H., Peinert, B., Borcherding, J., Leese, F. (2020): Umwelt-DNA besstätigt die Wiederbesiedlung und Ausbreitung der Rheingroppe in einem restaurierten Fließgewässer. Natur u Landsch. 95 (8): 358-363. doi 10.17433/8.2020.50153827.358-363

Tavares, C.N., Brauns, M., Hille, S., Krenek, S., Borcherding, J., Weitere, M. (2020): Tracing the colonization process of non-native gobies into a large river: the relevance of different dispersal modes. Biol Invasions 22: 2421–2429.


Fortes Silva, R., Heubel, K., Simon, M.-V., Borcherding, J. (2019): Have a break or keep going - behavioral and metabolic overwintering strategies of two invasive species of the River Rhine, Germany. Environ. Biol. Fish. 102, 1057-1068. DOI: 10.1007/s10641-019-00890-7

Borcherding, J., Heubel, K., Storm, S. (2019): Competition fluctuates across years and seasons in a 6-species-fish community: empirical evidence from the field. Rev. Fish biol. Fish. 29: 589-604

Økland, F., Havn, T.B., Thorstad, E.B., Heermann, L., Sæther, S.A., Tambets, M., Teichert, M. A.K.., Borcherding, J. (2019): Mortality of downstream migrating European eel at power stations can be low when turbine mortality is eliminated by protection measures and safe bypass routes are available Inter. Rev. Hydrobiol. 104: 68-79


Borcherding, J., Kiel, S., Struebig, A., Arndt, H. (2017): Vertical migration of Daphnia galeata – Evidence for the use of an alternative resource from a lethal environment. Limnologica 62: 132-139

Havn, T.B., , Økland, F., Teichert, M.A.K., Heermann, L. , Borcherding, J. , Sæther, S.A., Tambets, M., Diserud, O.H., Thorstad, E.B. (2017): Movements of dead fish in rivers. Anim. Biotelemetry (2017). DOI 10.1186/s40317-017-0122-2

Havn, T.B., Sæther, S.A., Thorstad, E.B., Teichert, M.A.K., Heermann, L., Diserud, O.H., Borcherding, J., Tambets, M., Økland, F. (2017) Downstream migration of Atlantic salmon smolts past a low head hydropower station equippped with Archimedes screw and Francis turbines. Ecological Engineering 105: 262–275

Havn, T.B.,  Thorstad, E.B., Teichert, M.A.K.,  Sæther, S.A., Heermann, L.,  Hedger, R.D., Tambets, M.  et al. (2017): Hydropower-related mortality and behaviour of Atlantic salmon smolts in the River Sieg, a German tributary to the Rhine. Hydrobiologia DOI 10.1007/s10750-017-3311-3

Heermann L., DeAngelis, D.L., Borcherding, J. (2017):A new mechanistic approach for the further development of a population with established size bimodality. PLOS ONE. DOI 10.1371/journal.pone.0179339

Heynen, M., Bunnefeld, N., Borcherding,J. (2017): Facing different predators: adaptiveness of behavioral and morphological traits under predation. Current Zoology: 1-9

Økland, F., Teichert,  M. A.K., Havn, T.B., Thorstad, E.B., Heermann, L., Sæther, S.A., Meelis Tambets, M., Borcherding, J. (2017): Downstream migration of European eel at three German hydropower stations. NINA Report 1355

Thorstad, E.B., Havn, T.B., Saether, S.A., Heermann, L., Teichert, M.A.K., Diserud, G.H., Tambets, M., Borcherding, J., Økland, F. (2017): Survival and behaviour of Atlantic salmon smolts passing a run-of-river hydropower facility with a movable bulb turbine. Fish. Manag. Ecol. 2017: 1-9. DOI: 10.1111/fme.12216


Adrian-Kalchauser, I., Hirsch, P.E., Behrmann-Godel, J, Guyen,A.N., Watzlawczyk, S., Gertzen,S., Borcherding, J., Burkhardt-Holm, P. (2016): The invasive bighead goby Ponticola kessleri displays large-scale genetic similarities and small-scale genetic differentiation in relation to shipping patterns. Molecular Ecology 25: 1925-1943

Borcherding, J., Arndt, H., Breiden, S., Brenner, K., Heermann, L., Höfer, S., Leistenschneider, C., Lindner, J., Staas, S., Gertzen, S. (2016): Drift of fish larvae and juveniles in the Lower Rhine before and after the goby invasion. Limnologica 59: 53–62

Gertzen, S., Fidler, A., Kreische, F., Kwabek, L., Schwamborn V., Borcherding, J., (2016): Reproductive strategies of three invasive Gobiidae co-occurring in the Lower Rhine (Germany). Limnologica - Ecology and Management of Inland Waters 56, 39-48

Hellström, G., Heynen, M., Borcherding, J., Magnhagen, C. (2016): Individual consistency and context dependence in group-sizepreference of Eurasian perch. Behav. Processes 133: 6-11

Heynen, M., Borcherding, J., Bunnefeld, N., Magnhagen, C. (2016): Plasticity and consistency of behavioural responses to predation risk in laboratory environments. J. Zool. 300: 228-235

Langkau, M.C., Clavés, D., Schmidt, M.B., Borcherding, J. (2016): Spawning behaviour of Allis shad Alosa alosa: new insights based on imaging sonor data. J. Fish Biology 88: 2263-2274

Økland, F., Teichert, M. A. K.,  Thorstad, E. B.,  Havn, T .B.,  Heermann, L., Sæther, S. A., Diserud, O. H., Tambets, M., Hedger, R. D., Borcherding, J. (2016): Downstream migration of Atlantic salmon smolt at three German hydropower stations. NINA Rapport;1203: (pdf)


Ondracková, M., Valová, Z., Hudcová, I., Michálková, V.,  Andrea Simková, A., Borcherding, J., Jurajda, P. (2015): Temporal effects on host-parasite associations in four naturalized goby species living in sympatry. Hydrobiologia 746: 233-243


Borcherding, J., Breukelaar, A.W., Winter, H.V., König, U. (2014): Spawning migration and larval drift of anadromous North Sea houting (Coregonus oxyrinchus) in the River IJssel, the Netherlands. Ecology of Freshwater Fish 23: 161-170

Dierking, J., Phelps, L., Praebel, K., Ramm, G., Prigge, E., Borcherding, J., Brunke, M. & Eizaguirre, C. (2014): Anthropogenic hybridization between endangered migratory and commercially harvested stationary whitefish taxa (Coregonus spp.). Evolutionary Applications 7: 1068-1083

Ghomi, M.R., von Elert, E., Borcherding, J. & Fink, P. (2014): Fatty acid composition and content of round goby (Neogobius melanostomus Pallas 1814) and monkey goby (Neogobius fluviatilis Pallas 1814), two invasive gobiid species in the lower Rhine River (Germany). Journal of Applied Ichthyology 30: 527-531

Ghomi, M.R., von Elert, E., Borcherding, J., Uhde, A. & Fink, P. (2014): Correlation between body size and fatty acid and essential amino acid composition of round goby (Neogobius melanostomus) and monkey goby (Neogobius fluviatilis) from the Rhine River (Germany). Biologia 69: 799-805

Goldenberg, S.U., Borcherding, J., Heynen, M. (2014): Balancing the response to predation - the effects of shoal size, predation risk and habituation on behaviour of juvenile perch. Behaviour Ecology and  Sociobiology: 68: 989-998

Heermann, L., Scharf, W., van der Velde, G., Borcherding, J. (2014): Does the use of alternative food resources induce cannibalism in a size-structured fish population? Ecology of Freshwater Fish 23: 129-140

Heynen, M., Rentrop, I., Borcherding, J. (2014): Age-matters-Experienced predation risk affects behavior and morphology of juvenile 0+ and 1+ perch. Limnologica 44: 32-39

Kureck, A., Bieg, R., Wendeler, R. & Borcherding, J. (2014): Fecundity of the mayfly Ephoron virgo (Olivier, 1791) (Ephemeroptera: Polymitarcyidae): A long-term study in the River Rhine. Limnologica 47: 1-6


Borcherding, J., Dolina, M., Heermann, L., Knutzen, P., Krüger, S., Matern, S., van Treeck, R. & Gertzen, S. (2013): Feeding and niche differentiation in three invasive gobies in the Lower Rhine, Germany. Limnologica 43: 49-58

Borcherding, J., Hertel, A. & Breiden, S. (2013): Activity and competitive behaviour of invasive Neogobius melanostomus and Ponticola kessleri (Gobiidae) from the River Rhine, Germany. Ethology Ecology & Evolution 25: 351-365

Heermann, L., Borcherding, J. (2013): Competition, predation, cannibalism: the development of young-of-the-year perch populations in ponds with bream or roach. J. Appl. Ichthyol. 29: 549-554

Heermann, L., Emmerich, M., Heynen, M., Dorow, M., König, U., Borcherding, J., Arlinghaus, R. (2013): Explaining recreational angling catch rates of Eurasian perch, Perca fluviatilis: the role of natural and fishing-related environmental factors. Fisheries Management and Ecology 20: 187-200

Huss, M., Persson, L., Borcherding, J., Heermann,L. (2013): Timing of the diet shift from zooplankton to macroinvertebrates and size at maturity determine whether normally piscivorous fish can persist in otherwise fishless lakes. Freshwater Biology 58: 1416-1424

Lindner, K., Cerwenka, A.F., Brandner, J., Gertzen, S., Borcherding, J., Geist, J., Schliwen, U.K. (2013): Brief Communication. First evidence for interspecific hybridization between invasive goby species Neogobius fluviatilis and Neogobius melanostomus (Teleostei: Gobiidae: Benthophilinae). Journal of Fish Biology 82:2128-2134

Scharbert, A., Borcherding, J. (2013): Relationships of hydrology and life-history strategies on the spatio-temporal habitat utilisation of fish in European temperate river floodplains. Ecological Indicators 29: 348-360


Fink, P., Kottsieper, A., Heynen, M. & Borcherding, J. (2012): Selective zooplanktivory of an invasive Ponto-Caspian mysid and possible consequences for the zooplankton community structure of invaded habitats. Aquatic Sciences 74: 191-202

Haertl, A., Cerwenka, A.F., Brandner, J., Borcherding, J., Geist, J., Schliewen, U.K. (2012): First record of Babka gymnotrachelus (KESSLER, 1857) from Germany. SPIXIANA 35 (1): 155-159

König, U., Borcherding, J. (2012): Preserving young-of-the-year Perca fluviatilis in ethanol, formalin, or in a frozen state and the consequences for measuring morphometrics. J.Appl.Ichthyol.(2012): 1–5

Langkau, M.C., Balk, H., Schmidt, M.B., Borcherding, J. (2012): Can acoustic shadows identify fish species? A novel application of imaging sonar data. Fish. Manag. Ecol. 19: 313-322

Magnhagen, C., Hellström, G., Borcherding, J., Heynen, M. (2012): Boldness in two perch populations – long-term differences and the effect of predation pressure. J. Anim. Ecol. 81: 1311-1318


Borcherding, J., Gertzen, S., Staas, S. (2011): First record of Pontian racer goby, Babka gymnotrachelus (Gobiidae: Teleostei), in the River Rhein, Germany. Journal of Applied Ichthyology, online version 27: 1-3

Borcherding, J., Staas, S., Krüger, S., Ondra?ková, M., Šlapanský, L., Jurajda, P. (2011): Non-native Gobiid species in the lower River Rhine (Germany): recent range extensions and densities. Journal of Applied Ichthyology 27: 1-3

Hellström, G., Heynen, M., Oosten, J., Borcherding, J., Magnhagen, C. (2011): The effect of group size on risk taking and social conformity in Eurasian perch. Ecology of Freshwater Fish 20: 499-502

Heynen, M., Heermann, L., Borcherding, J. (2011): Does the consumption of divergent resources influence risk taking behaviour in juvenile perch (Perca fluviatilis L.)? Ecology of Freshwater Fish 20: 1–4.


Borcherding, J., Beeck, P., DeAngelis, D. L., Scharf, W. R. (2010): Match or mismatch: the influence of phenology on size-dependent life history and divergence in population structure. Journal of Animal Ecology 79(5): 1101-1112

Heynen, M., Hellström, G., Magnhagen, C., Borcherding, J. (2010): Does morphological variation between young-of-the-year perch from two Swedish lakes depend on genetic differences? Ecology of Freshwater Fish 19: 163-169

Borcherding, J., Heynen, M., Jäger-Kleinicke, T., Winter, H.V., Eckmann, R. (2010): Re-establishment of the North Sea houting in the River Rhine. Fisheries Management and Ecology 17: 291-293

Vohmann, A., Borcherding, J., Kureck, A., Bij de Vaate, A., Arndt, H. & Weitere, M. (2010): Strong body mass decrease of the invasive clam Corbicula fluminea during summer. Biological Invasions 12: 53-64


Heermann, L., Eriksson, L.-O., Magnhagen, C., Borcherding, J. (2009): Size-dependent energy storage and winter mortality of perch. Ecology of Freshwater Fish 18: 560-571

Scharf, W.R., Heermann, L., König, U., Borcherding, J. (2009): Development of abundance and size structure of young-of-the-year perch populations using three methods. Fisheries Research 96: 77-87


Borcherding, J. & Staas, S. (2008): Local riverine fish communities as promotors for habitat restoration in the floodplain area of the Lower Rhine . 4th World Fisheries Congress, Vancouver, Canada, 49: 835-843

J. Borcherding, C. Pickhardt, H. V. Winter, and J. S. Becker. Migration history of North Sea houting (Coregonus oxyrinchus L.) caught in Lake IJsselmeer (The Netherlands) inferred from scale transects of 88Sr:44Ca ratios. Aquatic Science 70:47-56

Borcherding, J., Magnhagen, C. (2008): Food abundance affects both morphology and behaviour of juvenile perch. Ecology of Freshwater Fish 17: 207-218

Magnhagen, C., Borcherding, J. (2008): Risk-taking behaviour in foraging perch: does predation pressure influence age-specific boldness? Animal Behaviour 75: 509-517

Urbatzka, R., Beeck, P., van der Velde, G. & Borcherding, J. (2008): Alternative use of food resources causes intra-cohort variation in the size distribution of young-of-the-year perch  (Perca fluviatilis). Ecology of Freshwater Fish: 17 (3): 475-480


Klein Breteler, J.K., Vriese,T., Borcherding, J., Breukelaar, A., Jörgens, L., Staas, S., deLaak, G., Ingendahl, D. (2007): Assessment of population size and migration routes of silver eel in the River Rhine based on a 2-year combined mark-recapture and telemetry study. ICES Journal of Marine Science 64: 1450-1456

Heermann,L., Beek,P., Borcherding, J. (2007): Two size classes of 0+ year perch: is phenotiypic plasticity based on food resources? Journal of Fish Biology 70: 1365-1377

Borcherding J., Hermasch B., P.Murawski (2007): Field observations and laboratory experiments on growth and lipid content of young-of-the-year perch. Ecology of Freshwater Fish 16: 198-209


Borcherding, J., Murawski, S., Arndt, H. (2006). Population ecology, vertical migration and feeding of the Ponto-Caspian invader Hemimysis anomala in a gravel-pit lake connected to the River Rhine. Freshwater Biol. 51 (12): 2376-2387

Borcherding, J (2006): Prey or predator: 0+ perch (Perca fluviatilis) in the trade-off between food and shelter. Environmental Biology of Fishes 77 (1): 87-96

Borcherding, J., Scharbert, A. & Urbatzka, R. (2006). Timing of downstream migration and food uptake of juvenile North Sea houting stocked in the Lower Rhine and the Lippe (Germany). Journal of Fish Biology 68: 1271-1286

Borcherding, J.: (2006). Ten years of practical experience with the Dreissena-Monitor, a biological early warning system for continuous water quality monitoring. Hydrobiologia 556: 417-426

Heermann, L. & Borcherding, J. (2006). Winter short-distance migration of juvenile fish between two floodplain water bodies of the Lower River Rhine. Ecology of Freshwater Fish 15: 161-168


Borcherding, J., Sturm, W.: (2002). The seasonal succession of macroinvertebrates, in particular the zebra mussel (Dreissena polymorpha), in the River Rhine and two neighbouring gravel-pit lakes monitored using artificial substrates. International Revue of Hydrobiology 87: 165-181

Beeck, P.; Tauber, S.; Kiel, S.; Borcherding, J.: (2002). 0+ perch  predation on 0+ bream : a case study on a eutrophic gravel pit lake at the lower Rhine, Germany. Freshwater Biology 47: 2359-2369

Schöl, A., Kirchesch, V., Bergfeld, T., Schöll, F., Borcherding, J., Müller, D. (2002): Modelling the chlorophyll a content of the River Rhine – Interrelation between riverine algal production and population biomass of grazers, rotifers and the zebra mussel, Dreissena polymorpha. International Revue of Hydrobiology 87(2-3): 295-317

Weihrauch, F., Borcherding, J.: (2002). The zebra mussel, Dreissena polymorpha (Pallas) as an epizoon of anisopteran larvae (Anisoptera: Gomphidae, Corduliidae, Libelluidae). Odontologica 31(1): 85-94

Borcherding, J.; Bauerfeld, M.; Hintzen,D.; Neumann,D.: (2002). Lateral migrations of fishes between floodplain lakes and their drainage channels at the Lower Rhine: diel and seasonal aspects. Journal of Fish Biology 61: 1154-1170


Borcherding, J, & Wolf, J. (2001): The influence of suspended particles on the acute toxicity of 2-chlor-4-nitro-aniline, cadmium, and pentachlorophenol on the valve movement response of the zebra mussel ( Dreissena polymorpha). Archives of Environmental Contamination and Toxicology 40: 497-504


Borcherding, J., Maw, S.K. & Tauber, S. ( 2000): Growth of 0+ perch ( Perca fluviatilis ) predating on 0+ bream ( Abramis brama ). Ecology of Freshwater Fish 9: 236-241


Mulisch, M., Heep, T., Sturm, W. & Borcherding, J. (1998): Redescription of Ascobius lentus, a rare freshwater folliculinid (Ciliophora: Heterotrichida) from a pond in Germany. Acta Protozoologica 37: 29-39


Borcherding, J. & Jantz, B. (1997): The Dreissena-Monitor - The influence of pH and turbidity on the acute toxicity of pentachlorophenol under laboratory and in-situ conditions. Ecotoxicology 6:153-165


Borcherding, J. (1995): Laboratory experiments on the influence of food availability, temperature and photoperiod on gonad development in the freshwater mussel Dreissena polymorpha. Malacologia 36(1): 15-27


Borcherding, J. & Volpers, M. (1994): The "Dreissena-Monitor" - First results on the application of this biological early warning system in the continuous monitoring of water quality. In: "Proceedings of the International Conference on the Rehabilitation of the River Rhine", Arnhem, March 1993, J.A. van de Kraats (ed.), Wat. Sci. Tech. 29(3): 199-201

Kern, R., Borcherding, J. & Neumann, D. (1994): Recruitment of a freshwater mussel with a planktonic life-stage in running waters - studies on Dreissena polymorpha in the River Rhine. Arch. Hydrobiol. 131(4): 385-400


Borcherding, J. (1991): The annual reproductive cycle of the freshwater mussel Dreissena polymorpha Pallas in lakes. Oecologia 87: 208-218

Sprung, M. & Borcherding, J. (1991): Physiological and morphometric changes in Dreissena polymorpha (Mollusca; Bivalvia) during a starvation period. Malacologia 33: 179-191.

Review articles


Borcherding, J. 2011. The successful re-introduction of North Sea houting to the River Rhine System in Europe. In: Soorae, P. S. (ed.), Global Re-introduction Perspectives: 2011. More case studies from around the globe. Gland, Switzerland, IUCN/SSC Re-introduction Specialist Group and Abu Dhabi, pp 20-23.


Borcherding, J. 2010. Steps from ecological and ecotoxicological research to the monitoring for water quality using the zebra mussel in a biological early warning system. In: Van der Velde, G., Bij de Vaate, A., and Rajagopal, S. (eds.), The Zebra Mussel in Europe. Weikersheim, Margraf Publishers, pp 279-284.


Arndt, H., Borcherding, J., Kureck, A. (2002): The Ecology of Large Rivers – Preface. International Revue of Hydrobiology 87(2-3): 137


Neumann, D., & Borcherding, J. (1998): Die Fischfauna des Niederrheins und seiner ehemaligen Auenlandschaft – Istzustand, ökologische Anpassungen und Vorschläge für zukünftige Maßnahmen. LÖBF-Mitteilungen, Nr. 2/98: 12-14


Neumann, D., Borcherding, J. & Jantz, B. (1993): Growth and seasonal reproduction of Dreissena polymorpha in Rhine river and adjacent waters. - In: "Zebra mussels: biology, impacts, and control", T.F. Nalepa & D.W. Schloesser (eds.). Lewis Publishers, Boca Raton·Ann Arbor·London·Tokyo, pp 95-109

Applied research (selection)


Borcherding, J., Gertzen,S. (2016): Invasive Grundeln im Rhein. Eine Analyse nach sechs Jahren intensiver Forschung. Natur in NRW 2/16: 39-43

Borcherding, J., Gertzen, S. (2016): Die aktuelle Fischbestandsdynamik am Rhein unter besonderer Berücksichtigung invasiver Grundeln.  Ergebnisbericht zum Forschungsprojekt, Fischereiverbandes Nordrhein-Westfalen e.V. (Hrsg.), Münster, 48 Seiten


Borcherding, J. (2014): Der Nordseeschnäpel ist zurück im Rhein. Geglückte Wiederansiedlung einer ausgestorbenen Wanderfischart in NRW. Natur in NRW 4: 32-36

Kureck, A., Bieg, R., Wendeler, R., Borcherding, J. (2014): Fecunditiy of the mayfly Ephoron virgo (Olivier, 1791) (Ephemeroptera: Polymitarcidae): A long-term study in the River Rhine. Limnologica 47: 1-6


Borcherding, J. (2012): Invasive Grundeln – eine zweifelhafte „Erfolgsgeschichte“. DATZ 08: 34-37


Borcherding, J (2001):  Zwei neue Schwebgarnelen (Crustacea: Mysidacea) in rheinangebundenen Nebengewässern am Unteren Niederrhein. In: Beiträge zur Gewässer- und Naturkunde Nordrhein-Westfalens. S. Kronsbein, D. Pasch & K. van de Weyer (Hrsg.), Natur am Niederrhein (N.F.) 16 (1,2): 138-141


Borcherding, J., Barthold, K. & Becker, J. (1998): Entwicklungsnachweise für Brachytron pratense, Epitheca bimaculata, Libellula fulva und L. quadrimaculata (Odonata) in der Stopfenreuther Donauaue (Niederösterreich). Lauterbornia H. 33: 13-18

Borcherding, J. & Rehbach, R. (1998): Die Haffensche Landwehr – Ein Verbindungsgraben in der Rheinaue als überregional bedeutender Lebensraum heimischer Großmuscheln. LÖBF-Mitteilungen Nr. 2/98: 37-40

Bokranz, W., Neumann, D. & Borcherding, J. (1998): Der Komoran am Unteren Niederrhein – Untersuchungen zur Jagdstrategie und zum Beutefischspektrum. LÖBF-Mitteilungen, Nr. 2/98: 41-43

Hintzen, D., Bauerfeld, M. & Borcherding, J. (1998): Die Fischfauna in Grabensystemen des Deichhinterlands am Unteren Niederrhein - Ihre Bedeutung als Lebens- und Refugialraum für Fische. LÖBF-Mitteilungen Nr. 2/98, 31-34


Borcherding, J. (1997): Die Libellenfauna als Bioindikator für den Zustand einer Kulturlandschaft. LÖBF-Mitteilungen 2/97: 48-53

Engels, S. & Borcherding, J. (1997): Die Sanierungsmaßnahmen an der Kerspetalsperre (NRW): Langzeituntersuchungen über die Veränderungen des Makrozoobenthos im Ablauf der Talsperre. In: Tagungsbericht 1996 (Schwedt), Deutsche Gesellschaft für Limnologie (DGL), Krefeld, S. 226-230

Molls, F. & Borcherding, J. (1997): Der Fischegel Cystobranchus respirans (Troschel 1850) am Unteren Niederrhein mit neuen Daten zur Phänologie und zum Wirtsfischspektrum. Lauterbornia 28: 37-44


Borcherding, J. (1996): Möglichkeiten und Grenzen biologischer Frühwarnsysteme in der ökotoxikologischen Forschung und der kontinuierlichen Gewässerüberwachung dargestellt am Beispiel des Dreissena-Monitors. In: Tagungsbericht 1995 (Berlin), Deutsche Gesellschaft für Limnologie (DGL), Krefeld, S. 843-846

Faust, J. & Borcherding, J. (1996): Ökotoxikologische Untersuchungen mit dem Dreissena-Monitor - Zum Einfluß von Schwebstoffen auf die Toxizität verschiedener Substanzen. In: Tagungsbericht 1995 (Berlin), Deutsche Gesellschaft für Limnologie (DGL), Krefeld, S. 852-855


Diedrich, A., Neumann, D. & Borcherding, J. (1995): Flora und Fauna in Gräben einer niederrheinischen Auenlandschaft - Auswirkungen von Grabenräumungen. Natur und Landschaft 70(6): 263-268


Borcherding, J. (1994): The "Dreissena-Monitor" - improved evaluation of dynamic limits for the establishment of alarm-thresholds during toxicity tests and for continuous water control. In: Freshwater Field Test for Hazard Assessment of Chemicals. I.A. Hill, F. Heimbach, P. Leeuwangh & P. Matthiessen (eds.), Lewis Publishers, Boca Raton, pp 477-484

Borcherding, J., Becker, T., Schlegel, A. & Kureck, A. (1994): Beitrag zur Kenntnis der Odonatenfauna der Donauauen bei Stopfenreuth (Niederösterreich). Lauterbornia 15: 89-93

Borcherding, J., Kuhne, U., Lehmann-Greif, M., Ouali, S., Stahl, J. & Wipking, W. (1994): Zur Libellenfauna zweier Seen im Berner Oberland (Odonata). Ent. Ber. Luzern 31: 155-162

Borcherding, J. & Volpers, M. (1994): Der Dreissena-Monitor - Untersuchungen zum Verhalten der Zebramuscheln im Rhein - Störfallbewertung in der kontinuierlichen Gewässerüberwachung. Wasser und Boden, 8/1994, 46: 25-30

Borcherding, J. & Volpers, M. (1994): Der "Dreissena-Monitor" - Erfahrungen mit dem neuen biologischen Frühwarnsystem. In: "Biomonitore in der Gewässerüberwachung", Berlin, May 1993, H.-J. Pluta, J. Knie & R Leschber (Hrsg.), Schr.-Reihe Verein WaBoLu 93, Gustav-Fischer Verlag, Stuttgart·Jena·New York, pp 119-123


Borcherding, J. (1992): Morphometric changes in relation to the annual reproductive cycle in Dreissena polymorpha - a prerequisite for biomonitoring studies with zebra mussels. - In: "The zebra mussel Dreissena polymorpha - Ecology, Biological Monitoring and First Applications in the Water Quality Management", D. Neumann & H.A. Jenner (eds.). Gustav Fischer Verlag, Stuttgart·Jena·New York, pp 89-99

Borcherding, J. (1992): Another early warning system for the detection of toxic discharges in the aquatic environment based on the valve movements of the freshwater mussel Dreissena polymorpha. In: Limnologie aktuell - Vol. 4 - "The Zebra Mussel Dreissena polymorpha - Ecology, Biological Monitoring and First Applications in the Water Quality Management", D. Neumann & H.A. Jenner (eds.), Gustav Fischer Verlag, Stuttgart·Jena·New York, pp 127-146

Borcherding, J. (1992): Die Schalenbewegungen der Muschel Dreissena polymorpha als Monitorsystem zur Gewässerüberwachung. In: "Biologische Testverfahren. Beiträge zu den Biotest-Statusseminaren 1989 und 1992", K.G. Steinhäuser & P.-D. Hansen (eds.), Schr.-Reihe Verein WaBoLu 89, Gustav-Fischer Verlag, Stuttgart·Jena·New York, pp 361-373

Borcherding, J. & Ruyter Van Steveninck, E.D. DE (1992): Abundance and growth of Dreissena polymorpha larvae in the water column of the River Rhine during downstream transportation. - In: "The zebra mussel Dreissena polymorpha - Ecology, Biological Monitoring and First Applications in the Water Quality Management", D. Neumann & H.A. Jenner (eds.). Gustav Fischer Verlag, Stuttgart·Jena·New York, pp 29-44