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Borcherding, J., Aschemeier, D., Bruhy, J., Heermann, L., Lindner, J., Schröder, S. L., Wagner, K., Staas, S. (2021): The Caucasian dwarf goby, a new alien Gobiidae spreading at the Lower Rhine, Germany. J. Appl. Ichthyol. (pdf)  doi/10.1111/jai.14196

Schiwitza, S., Nitsche, F. (2020): A Needle in the Haystack – Mapping Sequences to Morphology Exemplified by the Loricate Choanoflagellate Enibas thessalia sp. nov. (Acanthoecida, Acanthoecidae). Protist 172 (pdf)

Wolter, C., Borcherding, J., Ferreira, T., Freyhof, J., Gessner, J., Górski, j., Nastase, A., Schomaker, C., Erös, T.. (2021): Characterization of European lampreys and fishes by their longitudinal and lateral distribution traits. Ecological Indicators 123. (pdf)


Arndt, H., Ritter, B., Rybarski, A., Schiwitza, S., Dunai, T., Nitsche, F. (2020) Mirroring the effect of geological evolution: Protist speciation in the Atacama Desert.  Global and Planetary Change (accept. April 15, 2020) (pdf)

Borcherding, J., Teichert, M., Heermann, L. (2020): Empfehlungen zum Schutz abwandernder Fische an Wasserkraftanlagen. Wasser u Abfall 09: 36-43 (pdf)

Borcherding, J., Webster, M.M., Heubel, K. (2020): Teaching laboratory for large cohorts of undergraduates: Private and social information in fish. Ecology and Evolution 10 (1): 4-10. (pdf)

Gooday, A.J., Schoenle, A., Dolan, J., Arndt, H. (2020). Protist diversity and function in the dark ocean - challenging the paradigms of deep-sea ecology with special emphasis on foraminiferans and naked protists. Europ. J. Protistol. 50, (pdf)

Havn, T.B., Thorstad, E.B., Borcherding, J., Heermann, L., Teichert, M.A.K., Ingendahl, D., Tambets, M., Sæther, S. A., Økland, F. (2020): Impacts of a weir and power station on downstream migrating Atlantic salmon smolts in a German river. River Research and Applications 36: 784-796. doi: 10.1002/rra.3590 (pdf)

Hempel, C. A., Jacobs, G., Beermann, A. J., Elbrecht, V., Macher, J.-N., Macher, T.-H., Peinert, B., Borcherding, J., Leese, F. (2020): Umwelt-DNA besstätigt die Wiederbesiedlung und Ausbreitung der Rheingroppe in einem restaurierten Fließgewässer. Natur u Landsch. 95 (8): 358-363. doi 10.17433/8.2020.50153827.358-363 (pdf)

Ingels, J., Vanreusel, A., Pape, E., Pasotti, F., Martínez Arbizu, P., Macheriotou, L., Vinther Sørensen, M., Edgcomb, V.P., Sharma, J., Sánchez, N., Homoky, W.B., Leduc, D., Gooday, A.J., Pawlowski, J., Dolan, J., Schratzberger, M., Gollner, S., Schoenle, A., Arndt, H., Woulds, C., Zeppilli, D. (2020) Ecological variables for deep-ocean monitoring must include microbiota and meiofauna for effective conservation. Nature Ecology & Evolution (in press) (pdf)

Sachs, M., Schluckebier, R., Poll, K., Schulz, V., Sabino-Pinto, J., Schmidt, E., Simon, K., Künzel, S., Ziegler, T., Arndt, H., Vences, M.  (2020): Evidence of Batrachochytrium dendrobatidis and other amphibian parasites in the Green toad (Bufotes viridis), syntopic amphibians and environment in the Cologne Bay, Germany. Salamandra 56  (in press) (pdf)

Schiwitza, S., Lisson, L., Arndt, H., Nitsche, F. (2020): Morphological and molecular investigation on freshwater choanoflagellates (Craspedida, Salpingoecidae) from the River Rhine at Cologne (Germany). Europ. J. Protistol. 73 (in press) (pdf)

Schoenle, A., Hohlfeld, M., Rosse, M., Filz, P., Nitsche, F., Arndt, H. (2020): Global comparison of bicosoecid Cafeteria-like flagellates from the deep ocean and surface waters, with reorganization of the family Cafeteriaceae. Europ. J. Protistol. 73  (in press) (pdf)

Tavares, C.N., Brauns, M., Hille, S., Krenek, S., Borcherding, J., Weitere, M. (2020): Tracing the colonization process of non-native gobies into a large river: the relevance of different dispersal modes. Biol Invasions 22: 2421–2429. (pdf)

Živaljić, S., Scherwass, A., Schoenle, A., Hohlfeld, M., Quintela-Alonso, P., Nitsche, F., Arndt, H. (2020): A barotolerant ciliate isolated from the abyssal deep-sea of the North Atlantic: Euplotes dominicanus sp. n. (Ciliophora, Euplotia). Europ. J. Protistol. 73  (in press)

Živaljić, S., Schoenle, A., Scherwass, A., Hohlfeld, M., Nitsche, F., Arndt, H. (2020): Influence of hydrostatic pressure on the behaviour of three ciliate species isolated from the deep sea. Marine Biology (in press) (pdf)



Borcherding, J., Heubel, K., Storm, S. (2019): Competition fluctuates across years and seasons in a 6-species-fish community: empirical evidence from the field. Rev. Fish biol. Fish. 29: 589-604 (pdf)

Fortes Silva, R., Heubel, K., Simon, M.-V., Borcherding, J. (2019): Have a break or keep going - behavioral and metabolic overwintering strategies of two invasive species of the River Rhine, Germany. Environ. Biol. Fish. 102, 1057-1068. DOI: 10.1007/s10641-019-00890-7 (pdf)

Økland, F., Havn, T.B., Thorstad, E.B., Heermann, L., Sæther, S.A., Tambets, M., Teichert, M. A.K.., Borcherding, J. (2019): Mortality of downstream migrating European eel at power stations can be low when turbine mortality is eliminated by protection measures and safe bypass routes are available. Inter. Rev. Hydrobiol. 104: 68-79 (pdf)

Schiwitza, S., Arndt, H., Nitsche, F. (2019) First description of an euryoecious acanthoecid choanoflagellate species, Enibas tolerabilis gen. et sp. nov., from a salar in the Chilean Andes based on morphological and transcriptomic data. Europ J. Protistol. 67: 106-113 (pdf)

Schoenle, A., Živaljić, S., Prausse, D., Voß, J., Jakobsen, K., Arndt, H. (2019): New phagotrophic euglenids from deep sea and surface waters of the Atlantic Ocean (Keelungia nitschei, Petalomonas acorensis, Ploeotia costaversata). Europ J. Protistol 69: 102-116  DOI: 10.1016/j.ejop.2019.02.007 (pdf)

van der Plas, F., Allan, E., Fischer, M., Alt, F., Arndt, H., Binkenstein, J., Blaser, S, Blüthgen, N., Böhm, S., Hölzel, N., Klaus, V.H., Kleinebecker, T., Morris, K., Oelmann, Y., Prati, D., Renner, S.C., Rillig, M.C., Schaefer, H.M., Schloter, M., Schmitt, B., Schöning, I., Schrumpf, M., Solly, E.F., Sorkau, E., Steckel, J., Steffan-Dewenter, I., Stempfhuber, B., Tschapka, M., Weiner, C.N., Weisser, W.W., Werner, M., Westphal, C., Wilcke, W., Manning, P. (2019): Towards the development of general rules describing landscape heterogeneity-multifunctionality relationships. Journal of Applied Ecology 56:168–179  DOI:10.1111/1365-2664.13260 (pdf)



Baker, P., Minzlaff, U., Schoenle, A., Schwabe, E., Hohlfeld, M., Jeuck, A., Brenke, N., Prausse, D., Rothenbeck, M., Brix, S., Frutos, I., Jörger, K.M., Neusser, T.P., Koppelmann, R., Devey, C., Brandt, A., Arndt, H. (2018): Potential contribution of surface-dwelling Sargassum algae to deep-sea ecosystems in the southern North Atlantic. Deep-Sea Res. II  148: 21-34 (pdf)

Heubel K (2018): Female mating competition alters female mating preferences in common gobies (Pomatoschistus microps). Current Zoology 64(3):351-361. Current Zoology, Volume 64, Issue 3, June 2018, Pages 351–361, (pdf)

de Jong K, Amorim MCP, Fonseca PJ, Heubel KU (2018): Noise affects multimodal communication during courtship in a marine fish. Frontiers in Ecology and Evolution 6(113). (pdf)

de Jong K, Amorim MCP, Fonseca PJ, Klein A, Fox CJ, Heubel KU (2018): Noise can affect acoustic communication and subsequent spawning success in fish. Environmental Pollution 237:814-823. (pdf)

Li, R., Nitsche, F. Arndt, H. (2018): Mesoscale investigations based on microsatellite analysis of the freshwater sponge Ephydatia fluviatilis in the River-Sieg system (Germany) reveal a genetic divergence. Conservation Genetics 19:959–968 (pdf)

Mück IM, Heubel, K (2018): Ecological variation along the salinity gradient in the Baltic Sea and its consequences for mating success in the common goby. Current Zoology 64(2):259-270. (pdf)

Schiwitza, S., Arndt, H., Nitsche, F. (2018): Four new choanoflagelle species from extreme saline environments: Indication for isolation-driven speciation exemplified by highly adapted Craspedida from salt flats in the Atacama Desert (Northern Chile). Europ. J. Protistol. 66: 86-96 (pdf)

Schoolmann, G., Arndt, H. (2018): Population dynamics of the invasive freshwater shrimp Neocaridina davidi in the thermally polluted Gillbach stream (North Rhine-Westphalia, Germany). Limnologica 71:1-7 (pdf)

Venter, P., Nitsche, F., Scherwass, A., Arndt, H. (2018): Discrepancies between molecular and morphological databases of soil ciliates studied for temperate grasslands of central Europe. Protist 169: 521-538 (pdf)

Venter, P., Nitsche, F., Arndt, H. (2018): The hidden diversity of flagellated protists in soil. Protist 169: 432-449 (pdf)

Weitere, M., Erken, M., Majdi, N., Arndt, H., Norf, H., Reinshagen, M., Traunspurger, W., Walterscheid, A. and Wey, J.K. (2018):  The food web perspective on aquatic biofilms. Ecological Monographs 88: 543–559; (pdf)

Živaljić, S.,  Schoenle, A., Nitsche, F., Hohlfeld, M., Piechocki, J., Reif, F., Shumo, M., Weiss, A., Werner, J., Witt, M., Voss, J., Arndt, H. (2018): Survival of marine heterotrophic flagellates isolated from the surface and the deep sea at high hydrostatic pressure: literature review and own experiments. Deep-Sea Res. II,Deep-Sea Res. II, 148: 251-259 (pdf)



Bleeker, K., de Jong, K., van Kessel, N., Hinde. C.A., Nagelkerke, L.A.J. (2017): Evidence for ontogenetically and morphologically distinct alternative reproductive tactics in the invasive Round Goby Neogobius melanostomus. PLOS ONE 12(4): e0174828. doi: 10.1371/journal.pone.0174828 (pdf)

Borcherding, J., Kiel, S., Struebig, A., Arndt, H. (2017): Vertical migration of Daphnia galeata – Evidence for the use of an alternative resource from a lethal environment. Limnologica 62: 132-139 (pdf)

Carr, M., Richter, D., Fozouni, P., Smith, T.J., Jeuck, A., Leadbeater, B., Nitsche, F. (2017): A six-gene phylogeny provides new insights into choanoflagellate evolution. Mol. Phylogenet. Evol. 107: 166-178 (pdf)

Groll, F., Arndt, H., Altland, A. (2017) Chaotic attractor in two-prey one-predator system originates from interplay of limit cycles. Theor Ecol 10 (2): 147–154 (pdf)

Havn, T.B., , Økland, F., Teichert, M.A.K., Heermann, L. , Borcherding, J. , Sæther, S.A., Tambets, M., Diserud, O.H., Thorstad, E.B. (2017): Movements of dead fish in rivers. Anim. Biotelemetry (2017). DOI 10.1186/s40317-017-0122-2 (pdf)

Havn, T.B., Sæther, S.A., Thorstad, E.B., Teichert, M.A.K., Heermann, L., Diserud, O.H., Borcherding, J., Tambets, M., Økland, F. (2017) Downstream migration of Atlantic salmon smolts past a low head hydropower station equippped with Archimedes screw and Francis turbines. Ecological Engineering 105: 262–275 (pdf)

Havn, T.B.,  Thorstad, E.B., Teichert, M.A.K.,  Sæther, S.A., Heermann, L.,  Hedger, R.D., Tambets, M.  et al. (2017): Hydropower-related mortality and behaviour of Atlantic salmon smolts in the River Sieg, a German tributary to the Rhine. Hydrobiologia DOI 10.1007/s10750-017-3311-3 (pdf)

Heermann L., DeAngelis, D.L., Borcherding, J. (2017):A new mechanistic approach for the further development of a population with established size bimodality. PLOS ONE. DOI 10.1371/journal.pone.0179339  (pdf)

Heynen, M., Bunnefeld, N., Borcherding,J. (2017): Facing different predators: adaptiveness of behavioral and morphological traits under predation. Current Zoology: 1-9 (pdf)

Jeuck, A., Nitsche, F., Wylezich, C., Wirth, O., Hennemann, M., Troll, N., Bergfeld, T., Monir, S., Scherwaß, A., Arndt, H. (2017): A comparison of quantification methods for heterotrophic flagellates of different taxonomic groups. Protist 168: 375-391 (pdf)

Økland, F., Teichert,  M. A.K., Havn, T.B., Thorstad, E.B., Heermann, L., Sæther, S.A., Meelis Tambets, M., Borcherding, J. (2017): Downstream migration of European eel at three German hydropower stations.NINA Report 1355 (pdf)

Richter, D., Nitsche, F. (2017): Choanoflagellata. in: Handbook of Protists. Archibald, J.M. Simpson, A.G.B , Slamovits, C.H.(eds.) Springer, 2nd edition ISBN-13: 978-3319281476

Ritz, S., Eßer, M., Arndt, H., Weitere, M. (2017) Large-scale patterns of biofilm-dwelling ciliate communities in a river network: only small effects of stream order. Intern. Rev. Hydrobiol. 102: 114-124 (pdf)

Schoenle, A., Nitsche, F., Werner, J., Arndt, H. (2017) Deep-sea ciliates: recorded diversity and experimental studies on pressure tolerance. Deep-Sea Res I, 128: 55-66 (pdf)

Schoolmann, G., Arndt, H. (2017):  Leaf-litter preferences of the invasive freshwater shrimp species Atyaephyra desmarestii and Neocaridina davidi. Crustaceana 90: 1715-1730 (pdf)

Thorstad, E.B., Havn, T.B., Saether, S.A., Heermann, L., Teichert, M.A.K., Diserud, G.H., Tambets, M., Borcherding, J., Økland, F. (2017): Survival and behaviour of Atlantic salmon smolts passing a run-of-river hydropower facility with a movable bulb turbine. Fish. Manag. Ecol. 2017: 1-9. (pdf)

Venter, P., Nitsche, F., Domonell, A., Heger, P., Arndt, H. (2017) The protistan microbiome of grassland soil: distribution in the mesoscale. Protist 168: 546-564 (pdf)

Vallon M & Heubel KU (2017): Egg density and salinity influence filial cannibalism in common gobies. Behavioral Ecology and Sociobiology 71:159. DOI 10.1007/s00265-017-2390-5 (pdf)



Adrian-Kalchauser, I., Hirsch, P.E., Behrmann-Godel, J, Guyen,A.N., Watzlawczyk, S., Gertzen,S., Borcherding, J., Burkhardt-Holm, P. (2016): The invasive bighead goby Ponticola kessleri displays large-scale genetic similarities and small-scale genetic differentiation in relation to shipping patterns. Molecular Ecology 25: 1925-1943 (pdf)

Arndt, H., Monsonís Nomdedeu, M. (2016): Protozoans and global climate change in aquatic systems, in: Marxsen, J. (Ed.), Climate Change and Microbial Ecology: Current Research and Future Trends. Caister Academic Press, Poole, UK, pp. 41-52 (pdf)

Borcherding, J., Arndt, H., Breiden, S., Brenner, K., Heermann, L., Höfer, S., Leistenschneider, C., Lindner, J., Staas, S., Gertzen, S. (2016): Drift of fish larvae and juveniles in the Lower Rhine before and after the goby invasion. Limnologica 59: 53–62 (pdf)

Blom EL, Mück I, Heubel K, and Svensson O (2016): Acoustic and visual courtship traits in two sympatric marine Gobiidae species - Pomatoschistus microps and Pomatoschistus minutus", Environmental Biology of Fishes 99(12): 999-1007. DOI: 10.1007/s10641-016-0550-5 (pdf)

Gertzen, S., Fidler, A., Kreische, F., Kwabek, L., Schwamborn V., Borcherding, J., (2016): Reproductive strategies of three invasive Gobiidae co-occurring in the Lower Rhine (Germany). Limnologica - Ecology and Management of Inland Waters 56, 39-48 (pdf)

Gossner, M.M., Lewinsohn, T., Kahl, T., Grassein, F., Boch, S., Prati, D., Birkhofer, K., Renner, S.C., Sikorski, J., Arndt, H., Baumgartner, V., Blaser, S., Blüthgen, N., Börschig, C., Buscot, F., Diekötter, T., Jorge, L.R., Jung, K., Keyel, A.C., Klein, A.-M., Klemmer, S., Krauss, J., Lange, M., Müller, J., Overmann, J., Pašalić, E., Penone, C., Perović, D., Purschke, O., Schall, P., Socher, S.A., Sonnemann, I., Tschapka, M., Tscharntke, T., Türke, M., Venter, P.C., Weiner, C.N., Werner, M., Wolters, V., Wurst, S., Westphal, C., Wubet, T., Fischer, M, Weisser, W.W., Allan, E. (2016): Land-use intensification causes homogenization of grassland communities. Nature 540: 266-269 (pdf)

Haileselasie, T.H., Mergeay, J., Weider, L.J., Sommaruga, R., Davidson, T.A., Meerhoff, M., Arndt, H., Jürgens, K., Jeppesen, E., De Meester, L. (2016): Environment not dispersal limitation drives clonal composition of arctic Daphnia in a recently deglaciated area. Molecular Ecology 25: 5830-5842 (pdf)

Hellström, G., Heynen, M., Borcherding, J., Magnhagen, C. (2016): Individual consistency and context dependence in group-sizepreference of Eurasian perch. Behav. Processes 133: 6-11 (pdf)

Heynen, M., Borcherding, J., Bunnefeld, N., Magnhagen, C. (2016): Plasticity and consistency of behavioural responses to predation risk in laboratory environments. J. Zool. 300: 228-235 (pdf)

Kalb N, Lindström K, Sprenger D, Anthes N, Heubel KU (2016): Male personality and female spawning consistency in a goby with exclusive male care. Behavioral Ecology and Sociobiology 70(5): 683-693. DOI: 10.1007/s00265-016-2089-z

Langkau, M.C., Clavés, D., Schmidt, M.B., Borcherding, J. (2016): Spawning behaviour of Allis shad Alosa alosa: new insights based on imaging sonor data. J. Fish Biology J. Fish Biology 88: 2263-2274 (pdf)

Nitsche, F. (2016): A phylogenetic and morphological re-investigation of Diaphanoeca spiralifurca, Didymoeca elongata and Polyoeca dichotoma(Acanthoecida/Choanomonadida) from the Caribbean Sea . European Journal of Protistology 52: 58–64 (pdf)

Nitsche, F., Thomsen, H. A., Richter, D.J. (2016): Bridging the gap between morphological species and molecular barcodes - Exemplified by loricate choanoflagellates. Europ. J. Protistol. 57: 26-37  (pdf)

Økland, F., Teichert, M. A. K.,  Thorstad, E. B.,  Havn, T .B.,  Heermann, L., Sæther, S. A., Diserud, O. H., Tambets, M., Hedger, R. D., Borcherding, J. (2016): Downstream migration of Atlantic salmon smolt at three German hydropower stations. NINA Rapport;1203: (pdf)

Pitsch, G., Adamec, L., Dirren, S., Nitsche, F., Šimek, K. Sirova, D., Posch, T. (2016) The green Tetrahymena utriculariae n. sp. (Ciliophora, Oligohymenophorea) with its Endosymbiotic Algae (Micractinium sp.), living in traps of a carnivorous aquatic plant. Protist 64: 322-335 (pdf)

Scherwass A., Erken M., Arndt H. (2016): Grazing effects of ciliates on microcolony formation in bacterial biofilms.In: Microbial Biofilms - Importance and Applications (edited by D. Dhanasekaran and N. Thajuddin), INTECH Open Scienc Publ., pp. 81-93, (ISBN 978-953-51-2436-8, Print ISBN 978-953-51-2435-1) (pdf)

Schönle, A., Jeuck, A., Nitsche, F., Venter, P., Prausse, D., Arndt, H. (2016): Methodological studies on estimates of abundance and diversity of heterotrophic flagellates from the deep-sea floor. J. Mar. Sci. Eng. 4, 22; doi:10.3390/jmse4010022 (pdf)

Schössow, M., Arndt, H., Becker, G. (2016): Response of gastropod grazers of the Lower River Rhine to food conditions, current velocity, and substratum roughness. Limnologica 58: 49-58 (pdf)

Soliveres, S., Manning, P., Prati, D., Gossner, M.M., Alt, F., Arndt, H., Baumgartner, V., Binkenstein, J., Birkhofer, K., Blaser, S., Bluthgen, N., Boch, S., Bohm, S., Borschig, C.,  Buscot, F., Diekotter, T., Heinze, J., Holzel, J., Jung, K., Klaus, V.H., Klein, A.-M.,  Kleinebecker, T., Klemmer, S., Krauss, J., Lange, M., Morris, E.-K., Müller, J., Oelmann, Y., Overmann, J., Pašalić, E., Renner, S.C., Rillig, M.C., Schaefer, H.M., Schloter, M., Schmitt, B., Schoning, I., Schrumpf, M., Sikorski, J., Socher, S.A., Solly, E.F., Sonnemann, I., Sorkau, E., Steckel, J., Steffen-Dewenter, I., Stempfhuber, B.,  Tschapka, M., Turke, M., Venter, P., Weiner, C.N., Weisser, W.W., Werner, M., Westphal, C., Wilcke, W., Wolters, V., Wubet, T., Wurst, S., Fischer, M., Allan, E. (2016): Locally rare species influence grassland ecosystem multifunctionality. Phil. Trans. R. Soc. B 371: 20150269. doi:10.1098/rstb.2015.0269 (pdf)

Soliveres, S., van der Plas, F., Manning, P., Prati, D., Gossner, M.M., Renner, S.C., Alt, F., Arndt, H., Baumgartner, V., Binkenstein, J., Birkhofer, K., Blaser, S., Blüthgen, N., Boch, S., Böhm, S., Börschig, C., Buscot, F., Diekötter, T., Heinze, J., Hölzel, N., Jung, K., Klaus, V.H., Kleinebecker, T., Klemmer, S., Krauss, J., Lange, M., Morris, E.K., Müller, J., Oelmann, Y., Overmann, J., Pašalić, E., Rillig, M.C., Schaefer, H.M., Schloter, M., Schmitt, B., Schöning, I., Schrumpf, M., Sikorski, J., Socher, S.A., Solly, E.F., Sonnemann, I., Sorkau, E., Steckel, J., Steffan-Dewenter, I., Stempfhuber, B., Tschapka, M., Türke, M., Venter, V., Weiner, C.N., Weisser, W.W., Werner, M., Westphal, C., Wilcke, W., Wolters, V., Wubet, T., Wurst, S., Fischer, M., Allan, E. (2016): Biodiversity at multiple trophic levels is needed for ecosystem multifunctionality. Nature 536: 456–459  doi:10.1038/nature19092 (pdf)

Thompsen, H.A., Nitsche, F, Richter, D.J. (2016): Seasonal occurrence of loricate choanoflagellates in Danish inner waters. Protist 167: 622-638 (pdf)

Weisse, T., Anderson, R., Arndt, H., Calbet, A., Hansen, P.J., Montagnes, D.J. (2016): Functional ecology of aquatic phagotrophic protists - concepts, limitations, and perspectives. Europ. J. Protistol. 55: 50–74 (pdf)


Berney, C., Geisen, S., Van Wichelen, J., Nitsche, F. Vanormelingen P., Bonkowski, M., Bass, D. (2015): Expansion of the ‘Reticulosphere’: Diversity of novel branching and network-forming amoebae helps to define Variosea (Amoebozoa). Protist 166: 271-295 doi: 10.1016/j.protis

Dunthorn, M., Lipps, J.R., Dolan, J.R. et al (2015): Ciliates - protists with complex morphologies and ambiguous early fossil record. Marine Micropaleontology (pdf)

Nitsche, F., Arndt, H. (2015) Comparison of similar Arctic and Antarctic morphotypes of heterotrophic protists regarding its genotypes and ecotypes. Protist 166: 42-57 (pdf)

Ondracková, M., Valová, Z., Hudcová, I., Michálková, V.,  Andrea Simková, A., Borcherding, J., Jurajda, P. (2015): Temporal effects on host-parasite associations in four naturalized goby species living in sympatry. Hydrobiologia 746: 233-243 (pdf)

Schoolmann, G., Nitsche, F., Arndt, H. (2015) Aspects of the life span and phenology of the invasive freshwater shrimp Atyaephyra desmarestii (Millet, 1831) at the northeastern edge of its range (Upper Rhine). Crustaceana 88: 949-962 (pdf)


Borcherding, J., Breukelaar, A.W., Winter, H.V., König, U. (2014): Spawning migration and larval drift of anadromous North Sea houting (Coregonus oxyrinchus) in the River IJssel, the Netherlands. Ecology of Freshwater Fish 23: 161-170 (pdf)

Deng, L., Krauss, S., Feichtmayer, J. Hofmann, R., Arndt, H., Griebler, C. (2014) Grazing of heterotrophic flagellates on viruses is driven by feeding behaviour. Environmental Microbiology Reports (2014) 6(4), 325–330, (pdf)

Dierking, J., Phelps, L., Praebel, K., Ramm, G., Prigge, E., Borcherding, J., Brunke, M. & Eizaguirre, C. (2014): Anthropogenic hybridization between endangered migratory and commercially harvested stationary whitefish taxa (Coregonus spp.). Evolutionary Applications 7: 1068-1083 (pdf)

Ghomi, M.R., von Elert, E., Borcherding, J. & Fink, P. (2014): Fatty acid composition and content of round goby (Neogobius melanostomus Pallas 1814) and monkey goby (Neogobius fluviatilis Pallas 1814), two invasive gobiid species in the lower Rhine River (Germany). Journal of Applied Ichthyology 30: 527-531 (pdf)

Ghomi, M.R., von Elert, E., Borcherding, J., Uhde, A. & Fink, P. (2014): Correlation between body size and fatty acid and essential amino acid composition of round goby (Neogobius melanostomus) and monkey goby (Neogobius fluviatilis) from the Rhine River (Germany). Biologia 69: 799-805 (pdf)

Goldenberg, S.U., Borcherding, J., Heynen, M. (2014): Balancing the response to predation - the effects of shoal size, predation risk and habituation on behaviour of juvenile perch. Behaviour Ecology and  Sociobiology: 68: 989-998 (pdf)

Heermann, L., Scharf, W., van der Velde, G., Borcherding, J. (2014): Does the use of alternative food resources induce cannibalism in a size-structured fish population? Ecology of Freshwater Fish 23: 129-140 (pdf)

Heynen, M., Rentrop, I., Borcherding, J. (2014): Age-matters-Experienced predation risk affects behavior and morphology of juvenile 0+ and 1+ perch. Limnologica 44: 32-39 (pdf)

Jeuck, A., Arndt, H., Nitsche, F. (2014): Extended phylogeny of the Craspedida (Choanomonada). Europ. J. Protistol. 50: 430-443 (pdf)

Kureck, A., Bieg, R., Wendeler, R., Borcherding, J. (2014): Fecunditiy of the mayfly Ephoron virgo (Olivier, 1791) (Ephemeroptera: Polymitarcidae): A long-term study in the River Rhine. Limnologica 47: 1-6 (pdf)

Marcus, H., Wey, j.K., Weitere, M. (2014): Disturbance alters the response of consumer communities towaards warming: a mesocosm study with biofilm-dwelling ciliates. Ecosphere 5(1): 1-15 (pdf)

Nitsche, F. (2014): Stephanoeca arndti spec. nov - First cultivation success including molecular and autecological data from a freshwater acanthoecid choanoflagellate from Samoa. Europ. J. Protistol. 50: 412-421

Wylezich,C., Kaufmann,D., Marcuse,M., Hülsmann,N. (2014): Dracomyxa pallida gen. et sp. nov.: a new giant freshwater foraminifer, with remarks on the taxon Reticulomyxidae (emend.). Protist 65, 854–869 (pdf)


Becks, L., Arndt, H. (2013): Different types of synchrony in chaotic and cyclic communities. Nature Communications 4: 1359, 1-9, DOI:10.1038 (pdf)

Borcherding, J., Dolina, M., Heermann, L., Knutzen, P., Krüger, S., Matern, S., van Treeck, R. & Gertzen, S. 2013. Feeding and niche differentiation in three invasive gobies in the Lower Rhine, Germany. Limnologica 43: 49-58 (pdf)

Borcherding, J., Hertel, A. & Breiden, S. (2013): Activity and competitive behaviour of invasive Neogobius melanostomus and Ponticola kessleri (Gobiidae) from the River Rhine, Germany. Ethology Ecology & Evolution 25: 351-365 (pdf)

Deng, L., Krauss, S., Feichtmayer, J. Hofmann, R., Arndt, H., Griebler, C. (2013): Grazing of heterotrophic flagellates on viruses is driven by feeding behaviour. Environmental Microbiology Reports, in press (pdf)

Domonell, A., Brabender, M., Nitsche, F., Bonkowski, M., Arndt, H. (2013): Community structure of cultivable protists in different grassland and forest soils of Thuringia. Pedobiologia 56: 1-7 (pdf)

Foissner, W., Wolf, K.W.,  Kumar, S., Xu, K, Quintela-Alonso, P. (2013): Five new spathidiids (Ciliophora: Haptorida) from Caribbean tank bromeliads. Acta Protozoologica 52(4)(pdf)

Heermann, L., Borcherding, J. (2013): Competition, predation, cannibalism: the development of young-of-the-year perch populations in ponds with bream or roach. J. Appl. Ichthyol. 29: 549-554 (pdf)

Heermann, L., Emmerich, M., Heynen, M., Dorow, M., König, U., Borcherding, J., Arlinghaus, R. (2013): Explaining recreational angling catch rates of Eurasian perch, Perca fluviatilis: the role of natural and fishing-related environmental factors. Fisheries Management and Ecology 20: 187-200 (pdf)

Huss, M., Persson, L., Borcherding, J., Heermann,L. (2013): Timing of the diet shift from zooplankton to macroinvertebrates and size at maturity determine whether normally piscivorous fish can persist in otherwise fishless lakes. Freshw. Biol. first published online : 27 MAR 2013, DOI: 10.1111/fwb.12138 (pdf)

Jeuck, A., Arndt, H. (2013): A short guide to common heterotrophic flagellates of freshwater habitats based on the morphology of living organisms. Protist 164: 842-860 (pdf)

Quintela-Alonso, P., Nitsche, F., Wylezich, C., Arndt, H., Foissner, W. (2013): A New Tetrahymena (Ciliophora, Oligohymenophorea) from Groundwater of Cape Town, South Africa.  J. Euk. Microbiol. 60: 235-246 (pdf)

Lindner, K., Cerwenka, A.F., Brandner, J., Gertzen, S., Borcherding, J., Geist, J., Schliwen, U.K. (2013): Brief Communication. First evidence for interspecific hybridization between invasive goby species Neogobius fluviatilis and Neogobius melanostomus (Teleostei: Gobiidae: Benthophilinae). J. Fish Biol. doi:10.1111/jfb.12127 (pdf)

Scharbert, A., Borcherding, J. (2013): Relationships of hydrology and life-history strategies on the spatio-temporal habitat utilisation of fish in European temperate river floodplains. Ecological Indicators 29: 348-360 (pdf)


Arndt, H. (2012): Protozoen. In: Grundwasserbiologie - Grundlagen und Anwendungen. DVGW-Information Wasser Nr. 75: 70-78 (pdf)

Brabender, M., Kiss, A.K., Domonell, A., Nitsche, F., Arndt, H. (2012): Phylogenetic and morphological diversity of novel soil cercomonad species with a description of two new genera (Nucleocercomonas and Metabolomonas). Protist 163: 495-528 (pdf)

Erken, M., Farrenschon, N., Speckmann, S., Arndt, H., Weitere, M. (2012): Quantification of individual fagellate - bacteria interactions within semi-natural biofilms. Protist 163: 632-642 (pdf)

Fink, P., Kottsieper, A., Heynen, M., Borcherding, J. (2012): Selective zooplanktivory of an invasive Ponto-Caspian mysid and possible consequences for the zooplankton community structure of invaded habitats. Aquat. Sci., 74: 191-202 (pdf)

Haertl, A., Cerwenka, A.F., Brandner, J., Borcherding, J., Geist, J., Schliewen, U.K. (2012): First record of Babka gymnotrachelus (KESSLER, 1857) from Germany. SPIXIANA 35 (1): 155-159 (pdf)

Langkau, M.C., Balk, H., Schmidt, M.B., Borcherding, J. (2012): Can acoustic shadows identify fish species? A novel application of imaging sonar data. Fish. Manag. Ecol. 19: 313-322 (pdf)

König, U., Borcherding, J. (2012): Preserving young-of-the-year Perca fluviatilis in ethanol, formalin, or in a frozen state and the consequences for measuring morphometrics. J.Appl.Ichthyol.: 1–5

Monsonís Nomdedeu, M., Willen, C., Schieffer, A., Arndt, H. (2012): Temperature dependent ranges of coexistence in a model of a two-prey-one-predator microbial food web. Marine Biology 159:2423–2430 (pdf)

Magnhagen, C., Hellström, G., Borcherding, J., Heynen, M. (2012): Boldness in two perch populations – long-term differences and the effect of predation pressure. J. Anim. Ecol. 81: 1311-1318 (pdf)

Pawlowski, J., Audic, S., Adl, S., Bass, D., Belbahri, L., Berney, C., Bowser, S.S., Cepicka, I., Decelle, J., Dunthorn, M., Fiore-Donno, A., Gile, H.G., Holzmann, M., Jahn, R., Jirk?, M., Keeling, J.P., Kostka, M., Kudryavtsev, A., Lara, E., Lukeš, J., Mann, G.D., Mitchell, A.D.E., Nitsche, F., Romeralo, M., Saunders, W.G., Simpson, G.B.A., Smirnov, V.A., Spouge, J., Stern, F.R., Stoeck, T., Zimmermann, J., Schindel, D., de Vargas, C. (2012): CBOL Protist Working Group: Barcoding eukaryotic richness beyond the animal, plant and fungal kingdoms. PLoS Biology 10 (11): e1001419 (pdf)

Risse-Buhl, U., Karsubke, M., Schlief, J., Baschien, C., Weitere, M., Mutz, M. (2012): Aquatic protists modulate the microbial activity associated with mineral surfaces and leaf litter. Aquatic Microbial Ecology, 66(2), 133-147 (pdf)

Salani, F.S.., Arndt, H., Hausmann, K., Nitsche, F., Scheckenbach, F. (2012): Analysis of the community structure of abyssal kinetoplastids revealed similar communities at larger spatial scales. ISME Journal 6: 713-723 (pdf)

Stoupin, D., Kiss, A.K., Arndt, H., Shatilovich, A.V., Gilichinsky, D.A., Nitsche, F. (2012): Cryptic diversity within the choanoflagellate morphospecies complex Codosiga botrytis – Phylogeny and morphology of ancient and modern isolates. Europ. J. Protistol. 48: 263-273 (pdf)

Viergutz, C., Linn, C., Weitere, M. (2012): Intra- and interannual variability surpasses direct temperature effects on the clearance rates of the invasive clam Corbicula fluminea. Mar. Biol. 159 (11): 2379-2387

Wey, J.K., Jürgens, K., Weitere, M. (2012): Seasonal and successional influences on bacterial community composition exceed that of protozoan grazing in river biofilms. Appl. Environ. Microbiol. 78 (6): 2013-2014 (pdf)


Ackermann, B., Esser, M., Scherwaß, A., Arndt, H. (2011): Long-term dynamics of microbial biofilm communities of the River Rhine with special references to ciliates. Int. Revue Hydrobiol. 96:1-19 (pdf)

Bergfeld, T., Scherwass, S., Ackermann, B., Fischer, H., Arndt, H., Schöl, A. (2011): Longitudinal and seasonal dynamics of the planktonic microbial community along the length of the River Rhine. River Systems 19(4): 337-349 (pdf)

Becker, G. (2011): Interactions between scrapers and periphyton.  In: Central European stream ecosystems: The long term study of the Breitenbach, Wagner, R., Marxsen, J., Zwick, P., Cox, E. (Eds), Wiley-VCH, Weinheim. ISBN 978-3-527-32952-6

Becks, L.; Agrawal, A.F. (2011): The effect of sex on the mean and variance of fitness in facultatively sexual rotifers. J. Evol. Biol. 24: 656-664 (pdf)

Borcherding, J., Gertzen, S., Staas, S. (2011): First record of Pontian racer goby, Babka gymnotrachelus (Gobiidae: Teleostei), in the River Rhein, Germany. J. Appl. Ichthyol. 27: 1399-1400 (pdf)

Borcherding, J., Staas, S., Krüger, S., Ondra?ková, M., Šlapanský, L., Jurajda, P. (2011): Non-native Gobiid species in the lower River Rhine (Germany): recent range extensions and densities. J. Appl. Ichthyol. 27: 1-3 (pdf)

Ellner, S.P.; Becks, L. (2011): Rapid prey evolution and the dynamics of two-predator food webs. Theoret. Ecol. 4: 133-152 (pdf)

Hellström, G., Heynen, M., Oosten, J., Borcherding, J., Magnhagen, C. (2011): The effect of group size on risk taking and social conformity in Eurasian perch. Ecol. Freshw. Fish 20: 499-502 (pdf)

Heynen, M., Heermann, L., Borcherding, J. (2011): Does the consumption of divergent resources influence risk taking behaviour in juvenile perch (Perca fluviatilis L.)? Ecol. Freshw. Fish 20: 1–4 (pdf) 

Howe, A.T., Bass, D., Scoble, J.M., Lewis, R., Vickerman, K., Arndt, H., Cavalier-Smith, T. (2011) Novel cultured protists identify deep-branching environmental DNA clades of Cercozoa: New genera Tremula, Micrometopion, Minimassisteria, Nudifila, Peregrinia. Protist 162: 332-372 (pdf)

Kaiser, T.S., Neumann, D., Heckel, D.G. (2011): Timing the tides: Genetic control of circdian and lunar phase of emergence is correlated in the marine midge Clunio marinus. BMC Genetics 2011: 12:49 (12pp.) (

Kathol, M., Fischer, H., Weitere, M. (2011): Contribution of biofilm-dwelling consumers to pelagic coupling in a large river. Freshw. Biol. 56: 1160-1172 (pdf)

Lagauzère S, Moreira S, Koschorreck M (2011): Influence of bioturbation on the biogeochemistry of the sediment in the littoral zone of an acidic mine pit lake. Biogeosci. 8: 339-352 (pdf)

Nitsche, F., Carr, M., Arndt, H., Leadbeater, B.S.C. (2011): Higher level taxonomy and molecular phylogenetics of the Choanoflagellatea. J. Eukaryot. Microbiol. 58: 452-462 (pdf)

Quintela-Alonso, P., Nitsche, F. Arndt, H. (2011): Molecular characterization and revised systematics of Microdiaphanosoma arcuatum (Ciliophora, Colpodea). J. Euk. Microbiol. 58: 114-119 (pdf)


Becks, L. and A.F. Agrawal (2010): Higher rates of sex evolve in spatially heterogeneous environments. Nature 468: 89-92 (pdf)

Borcherding, J., Beeck, P., DeAngelis, D. L., Scharf, W. R. (2010): Match or mismatch: the influence of phenology on size-dependent life history and divergence in population structure. J. Anim. Ecol. 79(5): 1101-1112 (pdf)

Borcherding, J., Heynen, M., Jäger-Kleinicke, T., Winter, H.V., Eckmann, R. (2010): Re-establishment of the North Sea houting in the River Rhine. Fisheries Manag. Ecol. 17: 291-293 (pdf)

Claessens, M., Wickham, S.A., Post, A.F., Reuter, M. (2010): A paradox of the ciliates? High ciliate diversity in a resource-poor environment. Mar. Biol. 157: 483-494 (pdf)

Heynen, M., Hellström, G., Magnhagen, C., Borcherding, J. (2010): Does morphological variation between young-of-the-year perch from two Swedish lakes depend on genetic differences? Ecol. Fresw. Fish 19: 163-169 (pdf)

Kaiser, T.S, Neumann, D., Hecke,l D.G, Berendonk, Th.U (2010): Strong genetic differentiation and postglacial origin of populations in the marine Midge Clunio marinus (Chironomidae, Diptera): Molecular Ecology 19: 2845-2857 (pdf)

Norf, H., Foissner, W. (2010): A new flagship peritrich (Ciliophora, Peritrichida) from the River Rhine, Germany. Apocarchesium arndti n. sp. J. Eukaryot. Microbiol. 57: 250-267 (pdf)

Norf, H., Kniggendorf, L.G., Fischer, A., Arndt, H., Kureck, A. (2010): Sexual and reproductive traits of Hypania invalida (Polychaeta, Ampharetidae): a remarkable invasive species in Central European waterways. Freshw. Biol. 55 (12): 2510-2520 (pdf)

Norf, H., Weitere, M. (2010): Resource quantity and seasonal background alter warming effects on communities of biofilm ciliates. FEMS microbiology ecology 74(2), 361-370 (pdf)

Scheckenbach, F., Hausmann, K., Wylezich, C., Weitere, M., Arndt, H. (2010): Large-scale patterns in biodiversity of microbial eukaryotes from the abyssal sea floor. PNAS 107(1): 115-120 (pdf)

Scherwass, A., T. Bergfeld, A. Schoel, M. Weitere und H. Arndt (2010): Changes in the plankton community along the length of the River Rhine: Lagrangian sampling during a spring situation. J. Plankt. Res. 32 (4): 491-502 (pdf)

Vohmann, A., Borcherding, J., Kureck, A., Bij de Vaate, A., Arndt, H., Weitere, M. (2010): Strong body mass decrease of the invasive clam Corbicula fluminea during summer. Biological Invasions 12: 53-64 (pdf)

Wylezich C., Nies G., Mylnikov A. P., Tautz D. & Arndt H. (2010): An evaluation of the use of LSU rRNA D1-D5 region for DNA based taxonomy of eukaryotic protists. Protist 161: 342-352 (pdf)


Bass, D., Chao, E. E.-Y., Nikolaev, S., Yabuki, A., Ishida, K., Berney, C., Pakzad, U., Wylezich, C., Cavalier-Smith, T. (2009): Phylogeny of Novel Naked Filose and Reticulose Cercozoa: Granofilosea cl. n.and Proteomyxidea Revised. Protist 160: 75-109 (pdf)

Bergfeld, T., Scherwass, A., Ackermann, B., Arndt, H., Schöl, A. (2009): Comparison of the components of the planktonic food web in three large rivers (Rhine, Moselle and Saar)". River Res. Appl. 25 (10): 1232-1250 (pdf)

Fleißner, G., Schuchhardt,K., Neumann, D., Bali, G., Falkenberg, G., Fleißner, G. (2008): A lunar clock changes shielding pigment transparency in larval ocelli of Clunio marinus. Chronobiology International 25: 17-30.

Heermann, L., Eriksson, L.-O., Magnhagen, C., Borcherding, J. (2009): Size-dependent energy storage and winter mortality of perch. Ecol. Freshw. Fish 18: 560-571 (pdf)

Kathol, M., Norf, H., Arndt, H., Weitere, M. (2009): Effects on temperature increase on the grazing of planctonic bacteria by biofilm-dwelling consumers. Aquat. Microb. Ecol. 55: 65-79 (pdf)

Loquay N., Wylezich C., Arndt H. (2009): Composition of groundwater nanoprotist communities in different aquifers based on aliquot cultivation and genotype assessment of cercomonads. Fund. Appl. Limnol. 174 (3):261-269 (pdf)

Neumann, D., Woermann, D. (2009): Physical conditions for trapping air by a microtrichia-covered insect cuticle during temporary submersion. Naturwissenschaften 96: 933-941.

Norf, H., Arndt, H., Weitere, M. (2009): Responses of biofilm-dwelling ciliate communities to planctonic and benthic resource enrichment. Microb. Ecol. 57: 678-700 (pdf)

Norf, H., Arndt, H., Weitere, M. (2009): Effects of resource supplements on mature ciliate biofilms: an empirical test using a new type of flow cell. Biofouling 25 (6): 769-778 (pdf)

Risse-Buhl, U., Scherwass, A., Schlüssel, A., Arndt, H., Kröwer, S., Küsel, K. (2009): Detachment and motility of surface-associated ciliates at increased flow velocities. Aquat. Microb. Ecol. 55: 209-218 (pdf)

Scharf, W.R., Heermann, L., König, U., Borcherding, J. (2009): Development of abundance and size structure of young-of-the-year perch populations using three methods. Fish. Res. 96: 77-87 (pdf)

Vohmann, A., Mutz, M., Arndt, H., Weitere, M. (2009): Grazing impact and phenology of the freshwater sponge Ephydatia muelleri and the bryozoans Plumatella emarginata and Fredericella sultana under experimental warming. Fresw. Biol. 54: 1078-1092 (pdf)

Weitere, M., Vohmann, A., Schulz, N., Linn, C., Dietrich, D. Arndt, H. (2009): Linking environmental warming to the fitness of the invasive clam Corbicula fluminea. Global Change Biol. 15: 2838-2851 (pdf)

Wey, J.K., Norf, H., Arndt, H., Weitere, M. (2009): Role of dispersal in shaping communities of ciliates and heterotrophic flagellates within riverine biofilms. Limnol. Oceanogr. 54(5): 1615-1626 (pdf)


Becks, L. and H. Arndt (2008): Transitions from stable equilibria to chaos, and back, in an experimental food web. Ecology 89 (11): 3222-3226 (pdf)

Borcherding, J., Magnhagen, C. (2008): Food abundance affects both morphology and behaviour of juvenile perch. Ecol. Freshw. Fish 17: 207-218 (pdf)

Borcherding, J., Pickhardt, C., Winter, H.V., Becker, J.S. (2008): Migration history of North Sea hounting (Corogonus oxyrinchus L.) caught in Lake Ijsselmeer (The Netherlands) inferred from scale transects of 88Sr:44Ca ratios. Aquat. Sci. 70: 47-56 (pdf)

Borcherding, J., Staas, S. (2008): Local riverine fish communities as promotors for habitat restoration in the floodplain area of the Lower Rhine . 4th World Fisheries Congress, Vancouver, Canada, 49: 835-843 (pdf)
Claessens, M., Wickham, S. A., Post, A.F., Reuter, M. (2008): Ciliate community in the oligotrophic Gulf of Aqaba, Red Sea. Aquat. Microbiol. Ecol. 53: 181-190

Magnhagen, C., Borcherding, J. (2008): Risk-taking behaviour in foraging perch: does predation pressure influence age-specific boldness? Anim. Behav. 75: 509-517 (pdf)

Nitsche, F. & Arndt, H. (2008): A new choanoflagellate species from Taiwan: Morphological and molecular biological studies of Diplotheca elongata nov. spec. and D. costata. Europ. J. Protistol. 44: 220-226 (pdf)

Urbatzka, R., Beeck, P., van der Velde, G., Borcherding, J. (2008): Alternative use of food resources causes intra-cohort variation in the size distribution of young-of-the-year perch  (Perca fluviatilis). Ecol. Freshw. Fish, 17(3): 475-480 (pdf)

Weitere, M., Dahlmann, J., Viergutz, C., Arndt, H. (2008): Differential grazer-mediated effects of high summer temperatures on pico- and nanoplankton communities. Limnol. Oceanogr. 53 (2): 477-486 (pdf)

Wey, J.K., Scherwass, A., Norf, H., Arndt, H., Weitere, M. (2008): Effects of protozoan grazing within river biofilms under semi-natural conditions. Aquat. Microb. Ecol. 52: 283-296 (pdf)


Borcherding J., Hermasch B., P. Murawski (2007): Field observations and laboratory experiments on growth and lipid content of young-of-the-year perch. Ecol. Freshw. Fish 16: 198-209.(pdf)

Cleven, E.-J., Königs, S. (2007): Growth of interstitial ciliates in association with ciliate bacterivory in a sandy hyporheic zone. Aquat. Microb. Ecol. 47: 177-189

Heermann, L., Beek, P., Borcherding, J. (2007): Two size classes of 0+ year perch: is phenotiypic plasticity based on food resources? J. Fish Biol. 70: 1365-1377 (pdf)

Klein Breteler, J.K., Vriese,T., Borcherding, J., Breukelaar, A., Jörgens, L., Staas, S., deLaak, G., Ingendahl, D. (2007): Assessment of population size and migration routes of silver eel in the River Rhine based on a 2-year combined mark-recapture and telemetry study. ICES J. Mar. Sci. 64: 1450-1456 (pdf)

Nitsche F., Weitere M., Scheckenbach F., Hausmann K., Wylezich C., Arndt H. (2007): Deep sea records of choanoflagellates with a description of two new species. Acta Protozool. 46 (2): 99-106 (pdf)

Norf, H., Arndt, H., Weitere, M. (2007): Impact of local temperature increase on the early development of biofilm-associated ciliate communities. Oecologia 151 (2): 341-350 (pdf)

Steinfartz, S. Weitere, M., Tautz, D. (2007): Tracing the first step to speciation: ecological and genetic differentiation of a salamander population in a small forest. Molec. Ecol. 16: 4550-4561

Viergutz, C., Kathol, M., Norf, H., Arndt, H., Weitere, M. (2007): Control of microbial communities by the macrofauna: a sensitive interaction in the context of extreme summer temperatures? Oecologia 151 (1): 115-124 (pdf)

Willkomm, M., Schlüssel, A., Reiz, E., Arndt, H. (2007): Effects of microcurrents in the boundary layer on the attachment of benthic heterotrophic nanoflagellates. Aquat. Microb. Ecol. 48: 169-174 (pdf)

Wylezich C., Mylnikov A. P., Weitere M., Arndt H. (2007): Distribution and phylogenetic relationships of freshwater Thaumatomonads with a description of the New Species Thaumatomonas coloniensis n. sp. J. Eukaryot. Microbiol. 54 (4): 347-357 (pdf)


Borcherding, J., Murawski, S., Arndt, H. (2006): Population ecology, vertical migration and feeding of the Ponto-Caspian invader Hemimysis anomala in a gravel-pit lake connected to the River Rhine. Freshw. Biol. 51 (12): 2376-2387 (pdf)

Borcherding, J. (2006): Prey or predator: 0+ perch (Perca fluviatilis) in the trade-off between food and shelter. Environ. Biol. of Fish. 77 (1): 87-96 (pdf)

Borcherding, J., Scharbert, A., Urbatzka, R. (2006): Timing of downstream migration and food uptake of juvenile North Sea houting stocked in the Lower Rhine and the Lippe (Germany). J. Fish Biol. 68: 1271-1286 (pdf)

Borcherding, J.. (2006): Ten years of practical experience with the Dreissena-Monitor, a biological early warning system for continuous water quality monitoring. Hydrobiologia 556: 417-426 (pdf)

Hausmann, K., Selchow, P., Scheckenbach, F., Weitere, M., Arndt, H. (2006): Cryptic Species in a Morphospecies Complex of Heterotrophic Flagellates: the Case Study of Caecitellus spp. Acta Protozool. 45: 415-431 (pdf)

Heermann, L. & Borcherding, J. (2006): Winter short-distance migration of juvenile fish between two floodplain water bodies of the Lower River Rhine. Ecol. Freshw. Fish 15: 161-168 (pdf)

Scheckenbach, F, Wylezich, C, Mylnikov, AP, Weitere, M, Arndt, H. (2006): Molecular comparisons of freshwater and marine isolates of the same morphospecies of heterotrophic flagellates. Appl Environ. Microb. 72 (10): 6638-6643 (pdf)


Becks, L., Hilker, F., Malchow, H., Jürgens, K., Arndt, H. (2005): Experimental demonstration of chaos in a microbial food web. Nature 435: 1226-1229 (pdf)

Fleissner, G., Schuchardt, K., Neumann,D., Fleissner, G. (2005): A moonlight receptor for circalunar rhythmicity in Clunio marinus? Shielding pigment in larval ocelli disappaers during moonlight nights. Proceedings European Biological Rhythm Society, III: 131

Fleissner, G., Schuchardt, K., Neumann,D., Fleissner,G. (2005): Organ-specific zeitgeber receptors: first candidate structures in the intestine of Clunio marinus. Proceedings Europeen Biological Rhythms Society III: 132.

Kreuzer, J., Auer, B., Scherwass, A., Arndt, H. (2005): Jahreszeitliche Sukzession und vertikale Verteilung des Zooplanktons im Fühlinger See. Ökologie und nachhaltige Entwicklung von Sport- und Freizeitseen 1: 77-94 (pdf)

Stauffer, D., Arndt, H. (2005): Simulation and experiment of extinction or adaptation of biological species after temperature changes. Int. J. Mod. Phys. C. 16 (3): 389-392 (pdf)

Scheckenbach, F., Wylezich, C., Weiter, M., Hausmann, K., Arndt, H. (2005). Molecular identity of strains of heterotrophic flagellates isolated from surface waters and deep-sea sediments of the South Atlantic based on SSU rDNA. Aquat. Microb. Ecol. 38 (3): 239-247 (pdf)

Scherwass, A., Fischer, Y., Arndt, H.(2005): Detritus as a potential food source for protozoans: consumption and digestion of plant detritus by a heterotrophic flagellate (Chilomonas paramecium) and a ciliate (Tetrahymena pyriformis). Aquat. Ecol.: 39 (4): 439-445  (pdf)

Scherwass, A., Arndt, H. (2005): Structure, dynamics and control of the ciliate fauna in the potamoplankton of the river Rhine. Archiv Hydrobiol: 164 (3): 287-307 (pdf)

Volpers, M., Neumann, D. (2005): Tolerance of two tubificid species (Tubifex tubifex and Limnodrilus hoffmeisteri) to hypoxic and sulfidic conditions in novel, long-term experiments. Arch. Hydrobiol. 164: 13-38 (2005)

Weitere, M., Scherwass, A., Sieben, K.-T., Friedrich, G., Arndt, H. (2005): Planctonic food web structure and potential carbon flow in the Lower River Rhine with a focus on the role of the Protozoans. River Res. Appl. 21: 535-549 (pdf)

Weitere, M., Bergfeld, T., Matz, C., Rice, S.A., Kjelleberg, S. (2005): Grazing resistance of Pseudomonas aeruginosa biofilms depends on type of protective mechanism, developmental stage and protozoan feeding mode. Environm. Microbiol. 7: 1593-1601 (pdf)


Auer,B., Czioska,E., Arndt, H. (2004): The pelagic community of a gravel pit lake: Significance of Coleps hirtus viridis (Prostomatida) and its role as a scavenger. Limnologica 34: 187-198

Auer, B., Elzer, U., Arndt, H. (2004): Comparison of pelagic food webs in lakes along a trophic gradient and with seasonal aspects: influence of resource and prdation. J. Plankt. Res. 26, 6: 697-709 (pdf)

Cleven, E.-J. (2004): Pelagic Ciliates in a Large Mesotrophic Lake: Seasonal succession and taxon-specific bacterivory in Lake Constance. Int. Revue Hydrobiol. 89 (3): 289-304 (pdf)

Cleven, E.-J. (2004): Seasonal and spatial distribution of ciliates in the sandy hyporheic zone of a lowland stream. Europ. J. Protistol. 40: 71-84 (pdf)

Dietrich, D., Arndt, H. (2004): Benthic heterotrophic flagellates in an Antarctic melt water stream. Hydrobiologia 529: 59-70 (pdf)

Matz, C., Deines, P., Boenigk, J., Arndt, H., Eberl, L., Kjelleberg, S., Jürgens, K. (2004): Impact of volacein-Ppoducing bacteria on survival and feeding of bacterivorous Nanoflagellates. Aquat. Microb. Ecol. 70 (3): 1593-1599 (pdf)

Stief, P., Altmann, D., de Beer, D., Bieg, R., Kureck, A. (2004): Microbial activities in the burrow environment of the potamal Ephoron virgo. Freshw. Biol. 49: 1152-1163 (pdf)

Weitere. M., Tautz, D. Neumann, D., Steinfartz, S. (2004): Adaptive divergence vs. environmental plasticity: tracing local genetic adaptation of metamorphosis traits in salamanders. Molec. Ecol. 13: 1665-1677 (pdf)

Weitere, M., Tautz, D. Neumann,D., und Steinfartz, S (2004).: Adaptative divergence versus environmental plasticity: tracing local genetic adaptation of metamorphosis traits in salamanders. Molecular Ecology 13: 1665-1677.

Wickham, S.A., Nagle, S., Hillebrand, H. (2004): Control of epibenthic ciliate communities by grazers and nutrients. Aquat. Microb. Ecol. 35: 153-162 (pdf)


Arndt, H., Hausmann, K., Wolf, M. (2003): Deep-sea heterotrophic nanoflagellates of the Eastern Mediterranean Sea: qualitative and quantitative aspects of their pelagic and benthic occurence. Mar. Ecol. Progr. Ser. 256: 45-56 (pdf)

Arndt, H.,  Schmidt-Denter, K., Auer, B., Weitere, M. (2003): Protozoans and biofilms. In: Fossil and Recent Biofilms (Krumbein, W.E., Paterson, D.M. & Zavarzin, G.A., Eds.) Kluwer Academic Publ., Dordrecht., p.173-189 (pdf)

Auer, B., Kiertucki, E., Arndt, H. (2003): Distribution of heterotrophic nano- and microflagellates in sediments of two small mesotrophic lakes. Archiv Hydrobiol. 158: 127-144 (pdf)

Cleven, E. and Meyer, E.I. (2003): A sandy hyporheic zone limited vertically by a solid boundary. Archiv Hydrobiol. 157: 267-288 (pdf)

Diederichs, S., Beardsley, C., Cleven, E.-J. (2003): Detection of ingested bacteria in benthic Ciliates using Fluorescence In Situ Hybridization. System. Appl. Microbiol. 26: 644-630 (pdf)

Garstecki, T. Wickham, S.A. (2003): The response of benthic rhizopods to sediment disturbance does not support the intermediate disturbance hypothesis. Oikos 103: 528-536

Neumann, D., Kappes, H. (2003): On the growth of bivalve gills initiated from a lobule producing budding zone. Bio. Bulletin 205: 73-82.

Prast, M., Arndt, H., Schöl, A. (2003): Untersuchungen zum planktischen Nahrungsnetz in Rhein und Mosel mittels Videomikroskopie. Hydrobiologie und Wasserbewirtschaftung 47: 102-107

Weitere, M., Arndt, H. (2003): Structure of the heterotrophic flagellate community in the water column of the River Rhine (Germany). Europ. J. Protistol. 39: 287-300

Weitere, M., Schmidt-Denter, K., Arndt, H. (2003): Laboratory experiments on the impact of biofilms on the plankton of a large river. Freshw. Biol. 48: 1983-1992 (pdf)


Beeck, P.; Tauber, S.; Kiel, S.; Borcherding, J. (2002): 0+ perch (Perca fluviatilis) predating on 0+ bream (Abramis brama): A case study on a eutrophic gravel pit lake at the lower Rhine, Germany. Freshw. Biol. 47: 2359-2369 (pdf)

Boenigk, J. (2002): Variability of ingestion rates with stage in cell cycle of a heterotrophic nanoflagellate (Spumella sp.) measured by an individual-based approach. Europ. J. Protistol. 38: 299-306 (pdf)

Boenigk, J., Arndt, H. (2002):  Bacterivory by heterotrophic flagellates: community structure and feeding strategies. Antonie van Leeuwenhoek 81: 465-480 (pdf)

Boenigk, J., Matz, C., Jürgens, K., Arndt, H. (2002): Food concentration-dependent regulation of food selectivity of interception-feeding bacterivorous nanoflagellates. Aquat. Microb. Ecol. 27: 195-202 (pdf)

Borcherding, J., Bauerfeld, M., Hintzen,D., Neumann, D. (2002): Lateral migrations of fishes between floodplain lakes and their drainage channels at the Lower Rhine: diel and seasonal aspects. J. Fish Biol. 61: 1154-1170 (pdf)

Borcherding, J., Sturm, W. (2002): The seasonal succession of macroinvertebrates in the River Rhine and two neighbouring gravel-pit lakes monitored with artificial substrates. Int. Revue Hydrobiol. 87: 165-181 (pdf)

Garstecki, T., Wickham, S.A., Arndt, H. (2002): Effects of experimental sediment resuspension on a coastal planktonic microbial food web. Estuarine, Coastal and Shelf Science 55: 751-762 (pdf)

Hausmann, K, Weitere, M., Wolf, M., Arndt, H. (2002): Meteora sporadica gen. nov. et sp. nov. (Protista incertae sedis) - an extraordinary free-living protist from the Mediterranean deep sea.  Europ. J. Protistol. 38: 171-177 (pdf)

Matz, C., Boenigk, J. Arndt, H., Jürgens, K. (2002): Role of bacterial phenotypic traits in selective feeding of the heterotrophic nanoflagellate Spumella sp. Aquat. Microb. Ecol. 27: 137-148  (pdf)

Neumann, D. (2002): Ecological rehabilitation of a degraded large river ystem – consideration based on case studies of macrozoobenthos and fish in the  Lower Rhineand ist catchment area. Int. Rev. Hydrobiology 87: 139-150.

Pfister, G., Auer, B., Arndt, H. (2002): Pelagic ciliates (Protozoa, Ciliophora) of different brackish and freshwater lakes – a community analysis at the species level.  Limnologica 32: 147-168 (pdf)

Scherwass A., Wickham, St.A., Arndt, H. (2002): Determination of the abundance of ciliates in highly turbid running waters - an improved method tested for the River Rhine. Arch. Hydrobiol. 156: 135-143 (pdf)

Schöl, A., Kirchesch, V., Bergfeld, T., Schöll, F., Borcherding, J., Müller, D. (2002): Modelling the chlorophyll a content of the River Rhine – Interrelation between riverine algal production and population biomass of grazers, rotifers and the zebra mussel, Dreissena polymorpha. Int. Revue Hydrobiol. 87 (2-3): 295-317 (pdf)

Stief, P., De Beer,D., Neumann, D. (2002): Small scale distribution of intertidal nitrite in freshwater sediment microcosms: the role of nitrate and oxygen availability, and sediment permeability. Microb. Ecology 43, 367 – 378.

Weihrauch, F., Borcherding, J. (2002): The zebra mussel, Dreissena polymorpha (Pallas) as an epizoon of anisopteran larvae (Anisoptera: Gomphidae, Corduliidae, Libelluidae). Odontologica 31 (1): 85-94 

Weitere, M., Arndt, H. (2002): Water discharge regulated bacteria/ heterotrophic nanoflagellates (HNF) interactions in the water column of the River Rhine. Microb. Ecol. 44:19-29 (pdf)

Weitere, M., Arndt, H. (2002): Top-down effects on pelagic heterotrophic nanoflagellates (HNF) in a large river (River Rhine): Do losses to the benthos play a role? Freshw. Biol. 47: 1437-1450 (pdf)


Adrian, R., Wickham, S. A., Butler, N. M. (2001):Trophic interactions between zooplankton and the microbial community in contrasting food webs: the epilimnion and deep chlorophyll maximum of a mesotrophic lake. Aquat. Microb. Ecol. 24: 83-97 (pdf)

Auer, B., Arndt, H. ( 2001): Taxonomic composition and biomass of heterotrophic flagellates in relation to lake trophy and season. Freshw. Biol. 46: 959-972 (pdf)

Boenigk, J., Matz, C., Jürgens, K., Arndt, H. (2001): The Influence of preculture conditions and food quality on the ingestion and digestion process of three species of Heterotrophic Nanoflagellates. Microb. Ecol. 42: 168-176 (pdf)

Boenigk, J., Arndt, H., Cleven, E.-J. (2001): The problematic nature of fluorescently labeled bacteria (FLB) in Spumella feeding experiments – an explanation by video microscopy. Archiv Hydrobiol. 152 (2): 329-338 (pdf)

Boenigk, J., Matz, C., Jürgens, K., Arndt, H.  (2001): Confusing selective feeding with differential digestion in bacterivorous nanoflagellates. J. Eukaryot. Microbiol. 48 (4): 425-432 (pdf)

Cleven, E.-J., Weisse, Th.  (2001): Seasonal succession and taxon-specific bacterial grazing rates of heterotrophic nanoflagellates in Lake Constance. Aquat. Microb. Ecol. 23: 147-161

Garstecki, T., Wickham, S.A. (2001): Effects of resuspension and mixing on population dynamics and trophic interactions in a simple benthic microbial food web. Aquat. Microb. Ecol. 25: 281-292 (pdf)

Ingendahl, D (2001): Dissolved oxygen concentration and emergence of sea trout fry from natural redds in tributaries of the River Rhine. J. Fish Biol. 58: 325-341

Neumann, D., Kramer,M., Raschke, I., Gräfe, B. (2001): Detrimental effects of nitrite on the development of benthic Chironomus larvae, in relation to their settlement in muddy sediments. Arch. Hydrobiol. 153: 103 – 128
Pfister, G. B., Auer, H., Arndt. (2001): Community analysis of pelagic ciliates in numerous different freshwater and brackish water habitats. Verh. Int. Verein. theor. angew. Limnol. 27: 3404-3408  (pdf)

Scherwass, A., Eimer, A., Arndt, H. (2001): Selective influence of filter-feeding benthic bivalves (Corbicula sp., Mytilus sp.) on planctonic ciliates. Verh. Int. Verein. theor. angew. Limnol. 27: 3315-3318 (pdf)


Arndt, H., Dietrich, D., Auer, B., Cleven, E., Gräfenhan, T., Weitere, M., Mylnikov, A.P.(2000): Functional diversity of hetrotrophic flagellates in aquatic ecosystems. In: The Flagellates (Leadbeater, B.S.C. & Green, J.C., Eds.) Taylor & Francis Ltd, London. 240-268 (pdf)

Boenigk, J. , Arndt, H. (2000): Comparative studies on the feeding behavior of two heterotrophic nanoflagellates: the filterfeeding choanoflagellate Monosiga ovata and the raprtorial-feeding kinetoplastid Rhynchomonas nasuta. Aquat. Microb. Ecol. 22: 243-249 (pdf)

Boenigk, J., Arndt, H. (2000): Particle handling during interception feeding by four species of heterotrophic nanoflagellate. J. Euk. Microbiol. 47: 350-358 (pdf)

Borcherding, J., Maw, S.K., Tauber, S. (2000): Growth of 0+ perch (Perca fluviatilis) predating on 0+ bream (Abramis brama). Ecol. Freshw. Fish 9: 236-241

Borcherding, J., Wolf, J. (2000): The influence of suspended particles on the acute toxicity of 2-chlor-4-nitro-aniline, cadmium, and pentachlorophenol on the valve movement response of the zebra mussel (Dreissena polymorpha). Arch. Environ. Con. Tox. 40: 497-504  

Dietrich, D., Arndt, H. (2000): Biomass partitioning of benthic microbes in a Baltic inlet: relationships between bacteria, algae, heterotrophic flagellates and ciliates. Mar. Biol. 136: 309-322 (pdf)

Garstecki, T., Arndt, H. (2000): Seasonal abundances and community structure of benthic Rhizopods in shallow lagoons of the Southern Baltic Sea. Europ. J. Protistiol. 36: 103-115

Garstecki, T., Verhoeven, R., Wickham, S.A., Arndt, H. (2000): Benthic-pelagic coupling: a comparison of benthic and planktonic heterotrophic protistan community structure in shallow estuaries of the Southern Baltic. Freshw. Biol. 45: 147-167 (pdf)

Premke, K., Arndt, H. (2000): Predation on heterotrophic flagellates by protists: Food selectivity determined using a live-staining technique. Archiv Hydrobiol. 150: 17-28 (pdf)

Wickham, S. A , Gieseke, A., Berninger, U.-G. (2000). Benthic ciliate identification and enumeration: an improved methodology and its application.  Aquat. Microb. Ecology 22: 79-91 (pdf)


Greve, G.D., van der Geest, H.G., Stuijfzand, S.C., Kureck, A., Kraak, M.H.S. (1999): Development and validation of an exotoxic test using field collected eggs of the riverine mayfly Ephoron virgo. Proc. Exper. & Appl. Entomol. 10: 105 - 110 (pdf)

Neumann, D. (1999): Aktuelle ökologische Probleme in Fließgewässern. 4. Franz-Ruttner Symposium ‘Lebensraum Fließgewässer’ Laufener Seminarbeiträge. Bayerische Akademie für Naturschutz und Landschaftspflege Lauffen.


Bartlett, F., Neumann, D. (1998): Sensitivity of Brown Trout alevins (Salmo trutta L. to nitrite at different chloride concentrations. Bull. Environ. Contam. Toxicol. 60: 340 - 346

Dietrich, D., Arndt, H. (1998): Untersuchungen zum benthischen und pelagischen mikrobiellen Nahrungsgewebe der Potter Cove, King George Island, Antarktische Halbinsel, unter besonderer Berücksichtigung der heterotrophen Flagellaten und Ciliaten. Ber. Polarforsch. 277: 25-26 (pdf)

Jantz, B., Neumann, D. (1998): Growth and reproductive cycle of the zebra mussel in the River Rhine as studied by a river bypass. Oecologia 114: 213 - 225
Pfister,G., Arndt, H. (1998): Food selectivity and feeding behaviour in omnivorous filter-feeding ciliates: A case study for Stylonychia. Europ. J. Protistol. 34: 446-457 (pdf)

Stief, P., Neumann, D. (1998): The formation of toxic nitrite concentrations in sediment cores of nitrate-loaded running waters. Archiv.Hydrobiol. 142: 153-169


Anlauf, A., Neumann, D. (1997): The genetic variability of Tubifex tubifex (Müller) in 20 populations and its relation to habitat type. Arch. Hydrobiol. 139: 145 - 162

Arndt, H. (1997): Spielen Protozoen bei der Manipulation pelagischer Nahrungsnetze eine Rolle? Laufener Seminararbeitr. 3/97: 77-88 (pdf)

Jürgens, K., Arndt, H, Zimmermann, H. (1997): Impact of metazoan and protozoan grazers on bacterial biomass distribution in microcosm experiments. Aquat. Microb. Ecol. 12: 131-138 (pdf)

Kahlert, M., Neumann, D. (1997): Early development of freshwater sponges under the influence of nitrite and pH. Arch. Hydrobiol. 139: 69 - 81

Neumann, D., Kaminsky, R., Heimbach, F. (1997): Timing of eclosion in marine insects on Mediterranean shores - studies on Clunio mediterraneus, C. ponticus and Thalassomyia frauenfeldi (Diptera: Chironomidae). Marine Biology 129: 513 - 521


Posch, T., Arndt, H. (1996): The uptake of sub-micrometre and µm-sized detrital particles by bacterivorous and omnivorous ciliates. Aquat. Microb. Ecol. 10: 45-53 (pdf)

Schiewer, U., Arndt, H., Heerkloss ,R., Jost, G., Walter, T., Wasmund, N. (1996): Experiments with controlled carbon flux variations in shallow brackish waters. Rostock. Meeresbiolog. Beitr. 4: 57-65 (pdf)


Arndt, H. & Berninger, U. (1995): Protists in aquatic food webs - complex interactions. In: G. BRUGEROLLE and J.-P. MIGNOT (eds.): Protistological Actualities, Clermont-Ferrand, France, p. 224-232
Mathes, J., Arndt, H. (1995): Annual cycle of protozooplankton (ciliates, flagellates and sarcodines) in relation to phyto- and metazooplankton in Lake Neumühler See (Mecklenburg, Germany). Arch. Hydrobiol. 134: 337-358 (pdf)


Arndt, H., (ed.)(1994): Protozooplankton Ecology - Methods and Seasonal Successions. Mar. Microb. Food Webs 8 (1 & 2): 1-375

Arndt, H. (1994): Protozoen als wesentliche Komponente pelagischer Ökosysteme von Seen. In: E. AESCHT (Hrsg.): Die Urtiere – eine verborgene Welt. Kataloge OÖ. Landesmuseums N.F. 71: 111-147
Salbrechter ,M.,  Arndt, H. (1994): The annual cycle of protozooplankton in the alpine, mesotrophic Lake Mondsee (Austria). Mar. Microb. Food Webs, 8: 217-234 (pdf)

Mathes, J., Arndt, H. (1994): Biomass and composition of protozooplankton in relation to lake trophy in north German lakes. Mar. Microb. Food Webs. 8: 357-375 (pdf)

Jürgens, K., Arndt, H., Rothaupt, K.O. (1994): Zooplankton-mediated changes of bacterial community structure. Microb. Ecol. 27: 27-42 (pdf)


Arndt, H. (1993): A critical review of the importance of rhizopods (naked and testate amoebae) and actinopods (Heliozoa) in lake plankton. Mar. Microb. Food Webs 7: 3-29 (pdf)

Arndt, H., Krocker, M., Nixdorf, B., Köhler, A. (1993): Long-term annual and seasonal changes of meta- and protozooplankton in Lake Müggelsee (Berlin): effects of eutrophication, grazing activities, and the impact of predation. Int. Revue ges. Hydrobiol. 78: 379-402 (pdf)

Nixdorf, B., Arndt, H. (1993): Seasonal changes in the plankton dynamics of a eutrophic lake including the microbial web. Int. Revue ges. Hydrobiol. 78: 403-410 (pdf)


Arndt, H., Güde, H., Macek, M., Rothaupt, K.O. (1992): Chemostats used to model the microbial food web: evidence for the feedback effect of herbivorous metazoans. Arch. Hydrobiol. Beih. Ergebn. Limnol. 37: 187-194 (pdf)


Arndt, H. (1991): On the importance of protozooplankton in the eutrophication process of the Baltic. Int. Revue ges. Hydrobiol. 76: 387-396

Arndt, H. (1991): Population dynamics and production of estuarine planktonic rotifers in the southern Baltic: Brachionus quadridentatus (HERMANN, 1783). Acta ichthyologica et piscatoria 21: 7-15 (pdf)

Arndt, H., Mathes, J. (1991): Large heterotrophic flagellates form a significant part of protozooplankton biomass in lakes and rivers. Ophelia 33: 225-234 (pdf)

Arndt, H., Nixdorf, B. (1991): Spring clear-water phase in a eutrophic lake: control by herbivorous zooplankton enhanced by grazing on components of the microbial web. Verh. Internat. Verein. Limnol. 24: 879-883 (pdf)

Nixdorf, B., Arndt, H., Schierhorn, E. (1991): Shortterm response within the microbial food web to organic matter loading. Acta ichthyologica et piscatoria S21: 163-170 (pdf)


Arndt, H. (1990): Das pelagische "microbial web" in einem eutrophen Flachsee: Jahreszeitliche Unterschiede in der Wechselwirkung zwischen Proto- und Matazooplankton. Deutsche Gesellschaft für Limnologie, Erweiterte Zusammenfassung der Jahrestagung 1990: 112-116 (pdf)

Arndt, H., Jost, G., Wasmund, N. (1990): Dynamics of pelagic ciliates in eutrophic estuarine waters: Importance of functional groups among ciliates and responses to bacterial and phytoplankton production. Arch. Hydrobiol. Beih. Ergebn. Limnol. 34: 239-245 (pdf)

Arndt, H., Schiewer, U., Jost, G., Wasmund, N., Walter, T., Heerkloss, R., Arlt, G., Arndt,E.-A. (1990): The importance of pelagic and benthic microfauna in a shallow-water community of the Darss-Zingst estuary, southern Baltic, during mesocosm experiments. Limnologica (Berlin) 20: 101-106 (pdf)

Arndt, H., Schröder, C., Schnese, W. (1990): Rotifers of the genus Synchaeta - an important component of the zooplankton in the coastal waters of the southern Baltic. Limnologica (Berlin) 21: 233-235 (pdf)

Schiewer, U., Arndt, H., Jost ,G., Heerkloss, R., Wasmund, N. (1990): Carbon flux dynamics in a shallow eutrophic estuary. Limnologica (Berlin) 20: 95-100 (pdf)


Arndt, H. (1989): Tageszeitliche Unterschiede in den Nahrungsketten pelagischer Ökosysteme. In: SCHÄLLER, G. & BÄHRMANN, R. (eds.)"Beiträge zur Ökophysiologie", Friedrich-Schiller-Universität Jena, p. 48-52
Arndt, H. (1989): Zooplankton production and its consumption by planktivores in a Baltic inlet. Proceedings of the 21st European Marine Biology Symposium, Gdansk, 14-19 September 1986, Polish Academy of Sciences - Institute of Oceanology, p. 205-214 (pdf)

Arndt,H., Heerkloss,R. (1989): Diurnal variation in feeding and assimilation rates of planktonic rotifers and its possible ecological significance. Int. Revue ges. Hydrobiol. 74: 261-272 (pdf)

Arndt, H., Waterstraat, A., Franke, D., Lenschow, U. (1989): Kurze Mitteilung über die Nahrung der kleinen Maräne (Coregonus albula L.) während eines Frühjahrsaspektes im Breiten Luzin (Mecklenburg). Fischeei-Forschung 27: 42-45 (pdf)


Arndt, H. (1988): Dynamics and production of a natural population of Brachionus plicatilis (Rotatoria, Monogononta) in a eutrophicated inner coastal water of the Baltic. Kieler Meeresforsch. Sonderh. 6: 147-153 (pdf)


Burckhardt, R., Arndt, H. (1987): Untersuchungen zur Konsumtion an Ciliaten durch Metazooplankter des Barther Boddens (südliche Ostsee). Meeersbiol. Beitr., WZ Rostock 36, math.-nat.R. 2: 22-26 (pdf)


Arndt, H. (1986): The relative importance of protozoans in a Baltic estuarine zooplankton community. Symp. Biol. Hung. 33: 257-261 (pdf)

Arndt, H., Schnese, W. (1986): Population dynamics and production of Acartia tonsa (Copepoda, Calanoida) in the Darss-Zingst estuary,  southern Baltic. Ophelia Suppl. 4: 329-334 (pdf)

Schiewer, U., Arndt, H., Baader, G., Börner, R., Evert, F.-K., Georgi,F., Heerkloss, R., Jost, G., Kell, V., Krüger, B., Walter, T.  (1986): The bounds and potential effects of NH4 (loading) on the pelagic system of a Baltic estuary. Limnologica (Berlin) 17: 7-28 (pdf)


Arndt, H. (1985): Eine Zählkammer für die mikroskopische Auswertung von Zooplanktonproben. WZ Rostock 34, math.-nat.R. 6: 30-31 (pdf)

Arndt, H., Kramer, H.-J., Heerkloss, R., Schröder, C. (1985): Zwei Methoden zur Bestimmung populationsdynamischer Parameter von Zooplanktern unter Laborbedingungen mit ersten Ergebnissen an Eurytemora affinis (Copepoda,Calanoida) und Synchaeta cecilia (Rotatoria, Monogononta). Meeresbiol. Beitr., WZ Rostock 34, math.-nat.R. 6: 17-21 (pdf)


Arndt, H., Debus L., Heerkloss, R., Schnese, W. (1984): Diurnal changes in the matter flux of a shallow-water ecosystem in a Baltic inlet. Ophelia Suppl. 3: 1-9 (pdf)

Arndt, H., Heerkloss, R., Schnese, W. (1984): Seasonal and spatial fluctuations of estuarine rotifers in a Baltic inlet. Limnologica 15: 377-385 (pdf)

Debus, L., Arndt, H. (1984): Nahrungsbiologische Untersuchungen an Jungfischpopulationen eines brackigen Flachwassergebietes des Barther Boddens (südliche Ostsee). Meeresbiol. Beitr., WZ Rostock 33, math.-nat.R. 6: 76-82 (pdf)

Heerkloss, R., Börner, R., Krummhaar, B., Arndt, H., Schnese, W. (1984): Growth inhibition in estuarine zooplankters due to the blue-green algae. Ophelia, Suppl. 3: 45-49 (pdf)

Heerkloss, R., Arndt, H., Hellwig, J., Vietinghoff, U., Georgi, F., WesselL, B., Schnese ,W. (1984): Consumption and assimilation by zooplankton related to primary production in the Baltic coastal inlet Barther Bodden. Limnologica 15: 387-394 (pdf)

Vietinghoff, U., Erdmann, N., Arndt, H., Kell, V., Hubert, M.-L. (1984): Integrated samples provide accurate means of parameters characterizing aquatic ecosystems. Int. Revue ges. Hydrobiol. 69: 121- 131 (pdf)

Vietinghoff, U., Hubert, M.-L., Heerkloss, R., Arndt, H., Schnese, W. (1984):  A mathematical submodel for the planktic Rotatoria in the ecosystem of the Barther Bodden. Int. Revue ges. Hydrobiol. 69: 159-172 (pdf)


Jansen, W., Arndt, H., Heerkloss, R. (1983): Die Rolle von Neomysis integer LEACH (Mysidacea, Crustacea) im Stoffumsatz des eutrophierten Brackwassergewässers Barther Bodden (südliche Ostsee). Wissenschaftliche Zeitschrift der Wilhelm-Pieck-Universität Rostock, Naturwissenschaftliche Reihe 5: 44-48 (pdf)


Arndt, H., Kobel, J., MIichalk, A., Wronna, F., Heerkloss, R.(1982): Untersuchungen zur Vertikalwanderung epibenthischer Copepoden mit Hilfe einer Planktonreuse. Meeresbiol. Beitr., WZ Rostock 31, math.-nat. R. 6: 57-60 (pdf)


Arndt, H., Schnese, W., Heerkloss, R.(1981): Untersuchungen zur räumlichen Verteilung des Zooplanktons in einem flachen inneren Küstengewässer der Ostsee. Meeresbiol. Beitr., WZ Rostock 30, math.-nat.R. 4/5: 43-48 (pdf)

Heerkloss, R., Arndt, H. (1981): Eine Lichtfalle zur Reinigung von Zooplanktonproben. Meeresbiol. Beitr., WZ Rostock 30, math.-nat.R. 4/5: 49-52 (pdf)


Heerkloss, R., Schnese, W., Arndt, H., Fischer, F. (1980): Konsumtionsrate und Vertikalwanderung des Zooplanktons in einem flachen inneren Küstengewässer der Ostsee. Meeresbiol. Beitr., WZ Rostock 29, math.-nat.R. 4/5: 73-76 (pdf)

Applied Research

Borcherding, J., Gertzen,S. (2016): Invasive Grundeln im Rhein. Eine Analyse nach sechs Jahren intensiver Forschung. Natur in NRW 2/16: 39-43. (pdf)

Borcherding, J., Gertzen, S., (2016): Die aktuelle Fischbestandsdynamik am Rhein unter besonderer Berücksichtigung invasiver Grundeln.  Ergebnisbericht zum Forschungsprojekt, Fischereiverbandes Nordrhein-Westfalen e.V. (Hrsg.), Münster, 48 Seiten. (pdf)

Borcherding, J. (2014): Der Nordseeschnäpel ist zurück im Rhein. Geglückte Wiederansiedlung einer ausgestorbenen Wanderfischart in NRW. Natur in NRW 4: 32-36. (pdf)

Borcherding, J. (2011): The successful re-introduction of North Sea houting to the River Rhine System in Europe. In: Soorae, P. S. (ed.), Global Re-introduction Perspectives: 2011. More case studies from around the globe. Gland, Switzerland, IUCN/SSC Re-introduction Specialist Group and Abu Dhabi, pp 20-23.

Borcherding, J. (2010): Steps from ecological and ecotoxicological research to the monitoring for water quality using the zebra mussel in a biological early warning system. In: Van der Velde, G., Bij de Vaate, A., and Rajagopal, S. (eds.), The Zebra Mussel in Europe. Weikersheim, Margraf Publishers, pp 279-284.

Umweltnetzwerk Erftstadt e.V., D. Neumann (Hg.) (2008): Natürliche Lebensgemeinschaften in der Börde. Fachtagung der Umweltakademie/Naturwerkstatt. Umweltzentrum Erftstadt, 51 S.

Ministerium für Umwelt, Naturschutz, Landwirtschaft und Verbraucherschutz (MUNLV) NRW (Hg) (2006): Leitfaden zur wasserwirtschaftlich-ökologischen Sanierung von Salmonidenlaichgewässern in NRW. Verfasser und Auftragnehmer: D. Neumann, St. Staas, A. Nemitz mit dem Aggerverband und dem Planungsbüro U. Koenzen, 166 S.

D. Neumann, S. Staas, Köln; U. Koenzen, Hilden; G. Mickoleit, R. Verhülsdonk, Gummersbach; F. Molls, Düsseldorf; A. Nemitz, Bonn; P. Podraza, Essen (2002): Ökologische Sanierung von Fließgewäassern und deren Interstitials – Pilotstudie an der Bröl aus Anlass der Lachswiedereinbürgerung im Siegsystem. Gewässerschutz, Wasser, Abwasser 188, 13/1-13/9.

Borcherding, J. (2001): Zwei neue Schwebgarnelen (Crustacea: Mysidacea) in rheinangebundenen Nebengewässern am Unteren Niederrhein. In: Beiträge zur Gewässer- und Naturkunde Nordrhein-Westfalens. S. Kronsbein, D. Pasch, van de Weyer; K. (Hrsg.), Natur am Niederrhein (N.F.) 16 (1,2): 138-141.

Bokranz, W. J. (2000): Beutespektrum des Kormorans: Bestimmungsschlüssel Otolithen und Knochenreste einheimischer Süßwasserfische. Ornithologische Mitteilungen 52. Jg. Nr. 10: 324-329.

Kureck, A., Bieg, R., Ottenberg, R. (2001): Einfluß von Futtermenge und Neozoen auf Überlebensrate und Wachstum  von Ephoron virgo (Ephemeroptera) im Rhein.  Deutsche Gesellschaft für Limnologie (DGL) - Tagungsbericht 2000 (Magdeburg): 379-83.

Kureck, A., Bieg, R. (2001): Zur Ernährungsökologie von Ephoron virgo (Ephemeroptera) im Rhein: Die Entwicklung der Mundwerkzeuge und der Einfluss von Nahrungskonkurrenz auf die filtierenden Larven.  Verhandlungen der Westdeutschen Entomologischen Tagung 2000: 299-306.

Huckenbeck, B., Wipking, W. (2000): (Über)-Leben in Raum und Zeit: Laufkäfer auf Biotopinseln in der Großstadt Köln. Mitt. Dtsch. Ges. Allg. Angew. Ent. 12, Giessen: 307-310.

Wolf, J., L. Küchler, Borcherding, J. (2000): Der Dreissena-Monitor: Ein Instrument zur kontinuierlichen Überwachung der Elbe an an der Gewässergütemeßstation Schmilka. 2. Hohenheimer Workshop zur Bioindikation "Biologische Testverfahren", Altbach-Deizisau, Deutschland, 6. – 8.4.1998; A. Fromin, U. Arndt, D. Elsner & A. Klump (Hrsg.), Verlag Günter Heimbach, Stuttgart.

D. Neumann (1999): Naturentwicklungsprojekte am Niederrhein - ein Beitrag zur nachhaltigen ökologischen Gestaltung des Niederrheins. Natur und Mensch Heft 5.

D.Neumann, D.Ingendahl, F.Molls, A.Nemitz (1998): Lachswiedereinbürgerung in NRW: Biologische Engpässe und Vorschläge für zukünftige Maßnahmen. LÖBF-Mitteilungen (Landesanstalt für Ökologie, Bodenordnung und Forsten, Nordrhein-Westfalen), Jahrgang 1998, Heft 2, 20-25.

D.Neumann in: Konzepte für eine Auenlandschaft am Unteren Niederrhein, unter besonderer Berücksichtigung der Gewässer. In: Naturschutzzentrum im Kreis Kleve e.V. (Hg.): Feuchtgebiete von internationaler Bedeutung, Schwerpunkt „Unterer Niederrhein“, S. 114-128, Rees-Bienen (1997).